List Of Madagascar (franchise) Characters - Madagascar Animals Movie

This is a list of characters that have appeared in the Madagascar films, the television series The Penguins of Madagascar, and the Netflix series All Hail King Julien.

Concept and creation

Tom McGrath explained in an interview that the intention of Madagascar was not to take a political stance on whether "zoos are bad and the wild is better, or that the wild is bad," but to show "the most extreme 'fish out of water' story that [they] could do". McGrath stated "the basic irony to the story is that, you think animals do belong in the wild, but if they're so accustomed to civility, they wouldn't know where food even came from," and the animals were meant to "love the zoo and to love where they are because they've got" 5,000 square feet (460 m²) "right off Fifth Avenue".

McGrath also described, during the research of Madagascar, they "found these crazy, weird animals that were already cartoons in their own right."

Main characters

Alex

Voiced by:

• Ben Stiller â€" in Madagascar franchise and specials
• Wally Wingert â€" in first video game and various spinoffs
• Quinn Dempsey Stiller/Declan Swift â€" as a cub
• Crispin Freeman â€" second and third video game

Alex (born Alakay) is an African lion. Alex is headstrong, protective and self-proclaimed leader of his friends. He is best friends with Marty (Zebra). He is used to the life of comfort in the zoo, but learns to adapt to the wild. He's smart, quick thinking and fast on his feet. He is a dancer by nature, often impressing others with his aerobatic feats. He does tend to be a little self-absorbed, often not seeing the problems of others above his own.

Alex enjoys his life at the zoo and has plenty of friends and is surprised when hearing Marty's wish to leave and live in the wild (Or run free in Connecticut as stated in the movie). As a cub, Alex was born on an African wildlife reserve and named Alakay. He has brown and yellow fur, whiskers, a black nose, a big mane, and a long tufted tail. He was the son of the alpha lion, Zuba, and was constantly given lessons on hunting, though he displayed no interest in the activity. During one of the sessions, Alex was captured by poachers, but the crate he was contained in fell into a river and eventually floated to New York. He is found by humans who renamed him Alex and took to the Central Park Zoo. In Madagascar, after Marty (Alex's best friend) leaves in an effort to take a train to the wild, Alex, Gloria, and Melman pursue him and are shot with tranquilizer darts by humans. The animal-rights activists pressure the zoo into releasing them back into the wild, but on the ship to the preserve, Alex and his friends fall into the ocean. They wash ashore on Madagascar, and, while Marty is excited to be free, Alex wishes to go home. He grows very hungry after days of being unable to eat, not realizing that steak is animal flesh. Hunger drives Alex into a crazed state, and after almost killing Marty in hunger (by biting him on the butt when he thought it was a piece of steak), in a moment of lucidity, he goes into self-imposed exile, barricading himself in the fossa territory of the island. Marty pursues him, unwilling to leave his friend, but is attacked by the fossa, the native predators of Madagascar. Alex works to defend Marty, and is later given sushi by the penguins to satiate his hunger.

In Madagascar: Escape 2 Africa, Alex encounters his parents, Zuba and Florrie, who are overjoyed to see their son and he becomes co-alpha lion alongside his father.

Zuba assumes Alex is strong after hearing he is the "king" of New York and arranges a rite of passage into the pride. Makunga, Zuba's rival, persuades Alex to challenge Teetsi during the Alex's rite of passage. Teetsi was revealed to be a bulky, strong lion and Alex is quickly defeated by him. To keep from banishing Alex, Zuba transfers his control to Makunga, and in turn, he, Florrie, and Alex are banished. Zuba soon grows unaccepting of Alex after realizing he is not a fierce warrior. The next day, the watering hole is shown to be dried up and Alex leaves with Marty to find out the cause. They leave the safety of the reserve, prompting Zuba to follow Alex and bring him back. Alex discovers that the stranded New Yorkers had created a dam, and through the use of dancing, he is recognized by the attacking New Yorkers. With the help of others, the dam is destroyed and they return to the preserve. His father learns to respect his habits, and they remain in Africa instead of flying home with the penguins. In The Penguins of Madagascar in a Christmas Caper, a stuffed animal Alex is seen briefly and its head is quickly dismembered by Nana's toy poodle, Mr. Chew. Alex has small cameos with his friends celebrating Christmas and singing a version of "Jingle Bells".

The fact that Alex loses the challenge to Teetsi and is spanked by Nana in Madagascar gives him the impression he is a horrible fighter, but this is not true, because (in the Nana case), she only gets Alex by surprise, but was more visible when in Madagascar 2, he has a short, but equilibrated fight against Nana. Alex is shown to have a superhuman level of interior physical strength, seeing that he broke the zoo's public telephone, and lifts Melman, Marty and Gloria with relative ease.

Alex made another appearance in "The Return of the Revenge of Dr. Blowhole." Wally Wingert reprises his role from the video games. In the show, he appeared as a hallucination "spirit guide" to help Skipper, whose memories had been stolen by the evil Dr. Blowhole. At first, Skipper thought he was very talkative and annoying, but soon he accepted his guidance, especially when he had to face Blowhole when he got back to New York City. A running gag in the episode is that Skipper's friends are confused and convinced he's crazy when they see him talk to Alex, since only Skipper can see him. After Dr. Blowhole was defeated, Alex's spirit guide form disappeared.

In Madagascar 3: Europe's Most Wanted, it opens on Alex's birthday and after a present from his friends of a mud New York he is determined to get the penguins from Monte Carlo. Alex creates four phases and after a fight with his friends they are exposed and are pursued by Captain Chantel DuBois.

After cutting the DuBois' snare they fly back to New York until they crash in a trainyard. Alex is determined to get back home and tries to fix the plane but fails. Then he persuades Gia to let them on the train, with whom he is smitten upon first sight. After being let on the train Alex and the gang realized that this was a ticket home.

After a failing performance in Rome he gives a pep talk and has the circus spiced up to get the circus promoter for a tour in America. Stefano told Alex why Vitaly was grouchy. Alex teaches Gia trapeze, and they begin to fall deeply in love. The London performance was better and they go to America. DuBois came to "take home" Alex but gets shot out of a cannon. Alex tells them why they had to join the circus.

Alex gives another pep talk to his friends and Julien that they were wrong and they decide to go back until DuBois comes. Awakening, Alex realizes they are back in the zoo with iron cage links between them. Luckily the circus came and rescued him. But DuBois is still determined to get Alex's head and is taking Stefano. But Alex saves the day (with Gia's help) by saving Stefano and defeating DuBois.

Marty

Voiced by:

• Chris Rock - in the films and specials
• Phil LaMarr - in the video games
• Thomas Stanley - as a foal

Marty is a plains zebra who used to live at the Central Park Zoo, and is a good friend of Alex. He once had an iPod before Alex accidentally broke it. Marty likes seaweed on a stick.

Marty is a hopeless optimist; he usually sees the brighter side of many things, taking every situation in stride. He has black and white stripes, a furry coat, a mane, and a tufted tail. He strives to be unique, which proves difficult when he meets his own kind in Africa, who look and act almost exactly the same. Marty is best friends with Alex and the two are rarely at odds. His carefree lifestyle tends to get on the nerves of his friends, but mostly serves to impress them; he is the one who helps hold the group together when things seem to get down.

Marty lived a life of comfort in the zoo. After his tenth birthday had come and gone, meaning that his life was already half over, Marty decided that never having seen the wild, he must travel to the wild, for at least a short time.

In the movie, Alex says he's black with white stripes, as he has 30 black, and only 29 white, but this was partly because Alex had to give an explanation as to why he was licking Marty, as he thought he was the steak in his dream.

The film begins on Marty’s 10th birthday (middle-aged for a zebra). His friends try to give him a great birthday, but somehow, Marty wants more. He escapes the Central Park Zoo, where he had lived for the past ten years, and makes a dash for the wild. His friends, Alex the Lion, Melman the Giraffe, and Gloria the Hippopotamus, flag him down, but as they are about to take him back to the zoo, police officers and animal control officers appear in all directions, tranquilize the animals, and send them on a ship to a wildlife preserve in Kenya. The penguins take over the ship and the four friends are washed up on the shores of Madagascar.

Once on the island, Marty had his dream fulfilled. Marty sets up a "bar" on the island, known as Casa del Wild (in English Home of the Wild). Alex is mad at Marty for getting them shipped to the wild, but he later apologizes to Marty and actually begins to enjoy himself on the island. Unfortunately, his enjoyment brings on his killer instincts, causing him to attack the resident lemurs and his friends, including Marty. Alex does not "go savage" for long, and is later able to rejoin his friends as they head back to the zoo.

Marty, along with his friends, has only a few small cameos in The Madagascar Penguins in a Christmas Caper. He, Alex, and Gloria are seen at the beginning decorating Melman's neck with Christmas lights. At the end, he, his friends, and almost all the other zoo animals are seen singing their own version of "Jingle Bells".

In Madagascar: Escape 2 Africa, Marty (along with Alex, Melman, Gloria, the lemurs, and the penguins) finds himself stranded in Africa after a failed attempt at flying to New York. There, he joins a herd of zebra that all look and sound exactly like him. While he enjoys this new lifestyle at first, he quickly becomes depressed when it becomes evident he has no individuality in a herd. Marty also becomes upset when Alex is unable to tell him apart from the other zebras. Ultimately, Alex is able to make amends by picking Marty out of the herd (later revealed to be due to the bite mark left on his backside from the previous film, and the fact that he is black with white stripes, while the other zebras are all white with black stripes.) Marty accompanies his friend to unblock a dam set by stranded New Yorkers in the jungle (which causes the watering hole to dry up). When Alex is caught, Marty gets the other animals to help rescue him.

In Madagascar 3 Europe's Most Wanted, Marty joins the circus along with the others. His circus act is cannon along with Stefano. In the middle of the film, you can tell he loves the afro. At the end of the film, it is shown that Marty is dancing in his afro (along with the others) to the tune Afro Circus / I like to Move it.

Melman Mankiewicz

Voiced by:

• David Schwimmer - in Madagascar films and specials
• Stephen Stanton - in Madagascar video games
• Zachary Gordon - as a calf

Melman Mankiewicz (Giacomo Manica) is a male reticulated giraffe. He has a furry coat, whiskers, a snout, a long neck, brown spots, horns, and a tufted tail. He is somewhat of a hypochondriac, having doses of medicine for meals and doctor's appointments frequently. He even complains about having brown spots on his neck (the spots, in real life, are natural).

He is skittish most of the time, but often has moments of profound clarity. His experience being medicated has given him insight regarding how to treat injured animals who might otherwise dig a hole and die in it. His love for Gloria is deep, and is always a gentleman to her; to Marty and Alex, he's just another one of the "awkward" guys.

Melman had a residence at the Central Park Zoo, where he received constant medical treatment for problems that were all psychosomatic. He would undergo MRIs, CAT scans, injections, flu shots, and be put into braces and crutches for no particular purpose at all. In the Central Park Zoo, Melman was used to his cushy life taking pills and undergoing treatments. One night, Marty the Zebra escaped and Melman immediately informed Alex of this.

In a flashback in Madagascar: Escape 2 Africa, there is a hint that he had a crush on Gloria as a child (as he seems uncomfortable when she comments that Alex is cute). When the animals nearly die trying to fly back to New York in a salvaged air plane, he screams he is in love with Gloria; she is asleep and does not hear. In Africa, Melman is horrified to learn that once a giraffe becomes sick, it has no way to survive and therefore must simply wait in a hole to die. He takes the position of Witch Doctor to cure sick animals, introducing them to Western medicine and giving them longer and better lives.

While he proves to be a very good witch doctor, he is informed by another giraffe that Melman has a brown spot on his shoulder, one which the previous Witch Doctor seemed to have gotten and "was gone" two days later. Thinking he only has two days to live, Melman initially waits in a hole, but then takes King Julien's advice and goes to declare his love to Gloria. He finds her on a date and simply tells her date what a wonderful girl Gloria is and to treat her right. When the watering hole dries up and King Julien decides to sacrifice an animal for the water gods, Melman offers himself. He is saved at the last moment, when Gloria tells him that she loves him back, and the two become a couple; it becomes apparent the "brown spot" was not fatal (the previous Witch Doctor is found alive and well, the brown spot in fact being a natural giraffe spot); however, there is not a scene where Melman is told of this. It is assumed that he figured it out when his "time" came and went and he was fine. He was last seen dancing off into the sunset with Gloria, Alex and Marty.

In Madagascar 3: Europe's Most Wanted, Melman and the other animals go to Monte Carlo to find the penguins. Once inside the hotel, Melman, Alex and Marty argue over who should be the leader of the group. They are crushed by Gloria's weight and they fall into the casino. Whilst escaping the hotel with the other animals, they are pursued by DuBois. On the way they lose DuBois and manage to reach the Hotel Ambassador. While boarding the plane, Melman is grabbed by the neck when DuBois appears. Alex with a little help from Mort, cuts the rope and saves Melman.

Above the skies of France, the plane's engine fails and it crash lands into a suburban rail yard. Whilst running from the authorities who are closing in on them, Melman gives Alex the idea of boarding the train containing the circus. Upon arrival in Rome, Melman and the group realize how terrible the circus was. En route to London, Melman and the others are discussing how terrible the circus was before Alex tells them about his newfound inspiration.

Melman and Gloria create a dancing tightrope act that is performed in London. When Melman and his friends are pursued by DuBois, the circus discover their true identities.

Gloria

Voiced by:

• Jada Pinkett Smith - in the films and specials
• Bettina Bush - in the video games
• Willow Smith - as a calf

Gloria is a hippopotamus who resided in the Central Park Zoo as an attraction, all the while being good friends with Alex and Marty, and Melman's girlfriend (as of Madagascar: Escape 2 Africa). She has gray skin, a snout, black fingernails, and a short stubby tail. She acts as the older sister to the gang, often putting her foot down when the boys start to lose control.

As a hippopotamus, Gloria is another animal in the Central Park Zoo and was one of the many attractions, drawing crowds that enjoyed seeing her swim about in her water tank. She would also receive the star treatment as one of the zoo's main attractions and attend Marty’s birthday party. At this party, Marty announced his wish upon blowing out the candles on his cake: to go to the wild. Gloria, Alex and Melman tried to have Marty come to his senses by telling him life out in the wilds was savage and difficult, whereas life in the zoos was adequate. Marty was unhappy at hearing this, and Gloria told Alex to go to console Marty. Later that night, Marty acted out his wish and headed straight for the Terminal Station to take a train and go to Connecticut, where Marty heard wide open spaces were found. Gloria urged both Alex and Melman to come with her to find Marty and bring him back before any people would notice. They were found, regardless, along with Mason and Phil, the two chimps, and a troupe of secretive penguins, all ending up tranquilized with darts.

When they all came to, they found themselves in crates bound for Africa. When the penguins hijack the ship, the crates fall into the ocean and wash up on the island of Madagascar. After some confusion, they start to set up their own society and attract the attention of a lemur colony.

In Madagascar: Escape 2 Africa, Gloria expresses a desire to join in the zoo breeding program, in the hopes of finding a boyfriend. When the animals crash land in Africa, she attracts the attention of a hippo named Moto Moto. Though she thinks he is the ideal boyfriend at first, she realizes that he only loves her for her large, "plumpy" body, where as Melman loves her for who she is. Realizing that she loves Melman back, she convinces him not to sacrifice himself into the volcano and Gloria grabs him by the horn and says, "It's crazy to think that I had to go half way around the world to find the perfect guy was right next door..." and then joins the other animals in rescuing Alex from Nana.

Gloria returns in Madagascar 3: Europe's Most Wanted. With Melman, the couple find their way as they contribute to the circus' success with a spectacular tightrope act.

The Penguins

Skipper

Voiced by:

• Tom McGrath

Skipper is the leader of the Adélie penguins (despite their color-schemes not matching the real species) and is the main character in the TV show. He has dark, sapphire-blue eyes and his body is wider than those of the other three penguins. Like the rest of the penguins, he has no idea what is expected of the penguin species, other than to "smile and wave." At one point, Private used a bird biology book to tell the other penguins how to act, but since the penguins usually need the chimpanzees Phil and Mason to translate for them, it was unclear how either he or Private were able to read the biology book, which is presumably written in English.

In the first "Madagascar" movie, Skipper is the mastermind behind a plot to escape the confines of the zoo and leave for Antarctica. Kowalski, Rico, and Private are his accomplices in this plot. They dig under their enclosure until they reach Marty the zebra, thinking it to be the South Pole. Skipper explains to Marty that they are attempting to escape the zoo because penguins do not belong there in captivity. He then orders his accomplices to continue tunneling. Late that night, Skipper and the other penguins are caught by a SWAT team, as are six other animals found outside the zoo. The penguins are tranquilized and put into a crate to be transferred.

When they awake, Skipper is confused at the lettering on the crate where they are being held and orders Mason to read it; Mason's companion Phil reads it. According to the box, they are all bound for a wildlife preserve in Africa, despite the cold climate penguins need. Skipper orders Rico to pick the lock on the crate where they are held, whereupon the four penguins knock out the crew and take command of the ship, commandeering it to reach Antarctica. Skipper and his mates finally reach Antarctica and are sorely disappointed by the barren, desolate terrain (as evidenced by Private's exclamation, "Well, this sucks...") Skipper has the ship return quickly from the South Pole, where it crashes into the beaches of Madagascar and a much warmer climate. Upon arriving, he and the other three encounter Gloria and Melman, and see that Alex and Marty (whom they refer to as their "monochromatic friend") are gone. Skipper decides to help rescue Marty from the grim fate of being eaten by the native foossa (possibly as gratitude for Marty’s silence regarding the escape). It is Skipper who helps to distract the foossa while Marty is recovered, and also helps fight off a number of them.

Skipper notices later on that Alex is still hungry, not having been able to eat steak, and has Rico prepare sushi for Alex. Skipper then relinquishes control of the ship to Alex, as he no longer needs it, and proceeds to sunbathe on the tropical beaches in comfort. Skipper is asked by Private if they should tell Alex and his friends that the ship is out of gas. Skipper declines, telling his troupe to "Just smile and wave, boys. Smile and wave."

In Madagascar 2, Skipper and the penguins salvage a crashed airplane to fly back to New York. They fix the plane when it crashes in Africa with the help of "more thumbs" (Mason and Phil). At the end of the movie, he marries a bobble head. In The Penguins of Madagascar, the bobble head is nowhere to be seen.

In Madagascar 3, it is revealed that Skipper, Kowalski, Rico and Private, along with Mason and Phil, have been in Monte Carlo. With Phil dressed as the "King of Versailles," they are able to amass a fortune of jewels, diamonds and gold and plan to buy an aircraft to pick up the "hippies" (Alex, Marty, Gloria, and Melman) in Africa and return to New York. Things change when Alex and the others rush to Monte Casino, believing that the penguins have abandoned them. Skipper later uses the fortune to help the gang become part of a traveling circus.

Skipper is one of the four main characters of The Penguins of Madagascar. In the show, many things are revealed about him and it expands on his character from the movies. In multiple episodes it is shown that Skipper knows karate and is an exceptionally capable fighter. He's also very strong, as seen in "Launchtime." In "Crown Fools," he states that he had taken down an angry walrus with "a wing and a prayer and another wing". However, in "Needle Point", it is revealed that he has a fear of needles, so in later episodes if he's injured he's treated with medical cream or the like. Generally, he is a straight man, displaying a very stereotypical general personality when not in battle: he's been shown waking his men up early in the morning for training, has given them trials on being able to work without him a number of times, and has been shown to being very hard-boiled. He is, though, soft on his men at times, properly rewarding them and even showing fear and worry when he knows they're in danger. He becomes especially upset when he believes one of them to be dead. He often tries to hide this part, as he once told Private that he (Private) is always bait because he "is gullible and expendable," but later becomes upset when Private is eaten.

Although Skipper's plans and theories often serve as jokes, his most unorthodox theories are sometimes proven true; the Blowhole existence is the most extreme example of this. Doctor Blowhole is first mentioned in "Eclipsed", when Skipper brings up his plan to extinguish the sun; this is because the sun was covered by a solar eclipse and, before Mason, Phil, and Kowalski mentioned it, Skipper believed he finally succeeded. Skipper being paranoid often comes up due to his theories, and whether or not he's right depends on the episode. Often, those around him refuse to follow along, mostly Marlene because she believes Skipper overreacts. He often comes up with theories without basis on new arrivals, usually flying piranha or being a spy for Doctor Blowhole. According to him, there are actually 8 continents, since he counts Atlantis (and appears to believe it is off the coast of Brazil). He once tells Maurice that if he had Skipper's security clearance, he would believe it too. Skipper indicates that he believes that some new arrivals to the zoo could be flying piranha. In the episode "Launchtime," when he asks Kowalski to search for a place where nearly no lemurs can be found, he states that he cannot set foot in Denmark; when Private questions him on this, he replies by saying "Well, that's private, Private, between me and the Danes." It is later revealed that the incident in question was caused by Hans the Puffin, revolving around a fish fight and top secret documents that made them both public enemy number one.

Skipper has mentioned that there used to be two more penguins (Manfredi and Johnson, named for two ill-fated characters in the World War II prisoner-of-war saga Stalag 17) in addition to himself and the other three; this is unconfirmed, and these claims have never been adequately explained. When they're brought up, it's usually as a warning or an aside remark, as they died in the past during a mission. In "The Hidden", he mentions that they died during a mission when they were attacked by "flying piranhas" and that they had to "bury what was left of them with a tea spoon". In "A Kipper for Skipper," when Skipper states that "relatively few penguins get left behind", as the others point out that Manfredi, Johnson, and another penguin were. In "Roger Dodger," he tells Rico that he would take him out if he had to and tells him "just ask Manfredi and Johnson." In "An Elephant Never Forgets," he tells them that while he was training Private that Manfredi and Johnson "fell for the exploding elephant foot." They are also mentioned by Private, who claims that they mistook the hind end of a beluga whale for an escape tunnel. In "Maurice at Peace," it is implied that they were accidentally smothered by the others due to a misinterpretation of a message which actually said to smother them with affection. In "Smotherly Love," it is mentioned that they died a very relaxing death in a day spa.

In The Penguins of Madagascar, he uses some Italian words such as "Sempre all'erta" (in "Haunted Habitat"), "Bravissimo" (in "The Hidden"), "Attenzione" (in "Crown Fools"), "Eccellente" (in "Kingdom Come"), "Finito!" (in "Popcorn Panic") and "Perfetto" (in "An Elephant Never Forgets"). His Italian pronunciation is not perfect. In "The Falcon and the Snow Job," Skipper falls for a female Peregrine falcon named Kitka and they date for a while. Skipper breaks up with her after it is revealed that she had swallowed Fred the squirrel, and in subsequent episodes she wasn't heard from or mentioned.

Skipper shares a very strong friendship with Marlene, a female Asian Otter who lives across from the penguins and is a main character in the series. Despite their conflicting differences and clashes of personality, the two still remain very close and are able to keep their bond of friendship strong. Marlene is also considered "privileged" by Skipper, due to the fact that he thinks of her as a smart, which is rather unusual, considering his otherwise generally scornful opinion of mammals. He sometimes allows her to tag along on missions.

As the series The Penguins of Madagascar continues, Skipper starts to realize that Private is following in his footsteps in "Private and the Winky-Factory." Skipper also becomes less sensitive to his enemies Dr. Blowhole and Hans the Puffin. Over the course of the series he likes and trusts them less and less. In the Dr. Blowhole Special "Blowhole Strikes Back," Hans and Skipper are referred to as "frenemies." At the beginning of the episode, Hans surprises Skipper by meeting him in China, and refers to him as such.

Kowalski

Voiced by:

• Jeff Bennett - in The Penguins of Madagascar
• Chris Miller - in the movies and other media

The tallest of the penguins, Kowalski is second-in-command, forms plans for the group and usually takes notes for Skipper. When Skipper is absent, he takes charge and his rank is First Lieutenant. He has blue eyes and tends to over-analyze situations.

Often formulating plans for the group, Kowalski is a loyal supporter of his leader, Skipper. Kowalski had escaped the zoo with the other penguins, but was promptly captured and put into a crate for transfer with his comrades. The crates were all placed on a ship bound for Africa, where all the creatures from the zoo would be kept in a wildlife preserve. While in the crate, Kowalski confessed that he could not read (he said the words on the side of the box were an "older code", in slight reference to Return of the Jedi) and asked another transferee, a chimpanzee named Phil, to read the words on the crate. As soon as this was done, the four penguins headed to the ship's bridge, knocked out the crew and captain, bound them up, and Kowalski plotted the ship's course to Antarctica.

He is the technician of the group and always has his notepad (and an abacus in the animated series). While being able to formulate plans and invent things (such as a rocket ship out of a trashcan and fireworks, a futuristic rotary saw blade made of energy and multiple random objects, or an intelligence sucking machine from just a cardboard box, a magnet, and a vacuum) he is shown to have a bit of difficulty deducing simpler machines. At the end of "Needle Point", it is revealed that he has an irrational fear of dentists, which is peculiar in that, as even Private soon states after Kowalski runs off, penguins do not have teeth. He seems to also spout nearly or completely nonsensical words, such as, "The moo cow may have a chocolate marshmallow", or "I'll be a bicycle cream cone", whenever either hit incredibly hard (as he did when body checked by a giant rat during a hockey game) or when shocked severely (as by Officer X's stun gun). He screamed "Galileo Galilei" when he got his injection during "Needle Point". He also loves candy.

In the series he is shown to be intelligent but it is not as apparent in the films (e.g. in Madagascar he does not know how to read and in Madagascar: Escape 2 Africa he states that the paper plane would help them fly back to NY and he does not understand that the fuel indicator was indicating that fuel is finished). He does most of his calculations on an abacus, although he was briefly in possession of a calculator that was destroyed. He is in love with a dolphin called Doris and recorded a poem for her on DVD which highlighted her apparent non-interest in him in the "Hot Ice" episode. (She was first mentioned in "Gone in a Flash" with a photo being shown.)

It has been noted that his inventions have a habit of horribly backfiring regardless of his best intentions. Some of his most significant blunders include: a time machine (or 'Chronotron') that triggered a temporal rift that nearly destroyed reality and resulted in three of Kowalski appearing in one place (noticeably, the first Kowalski from the future wanted to stop the ray and the second one wanted it to keep working); a ray designed to return spoiled food to its fresher state that turned Skipper into a chick; a mind-switching machine (which backfired in its first use because he forgot to take the warm-blooded/cold-blooded conflict into account when switching Rico with Roger); a device called the "Chronocurbulator" that stops time that left time seemingly permanently frozen when it broke down; a helmet that granted the user the power of telekinesis; an invisibility ray (or 'Transparent Matter-Maker', called 'Transmatterer' for short due to copyright issues) that turned itself invisible before being set to randomly fire; a mind-reading machine that quickly burnt out due to lack of processing power; a cloning machine that relied on the Higgs Boson particle for power; nanites that could reprogram other machines (these were programmed to not harm a penguin, but this backfired when they locked the penguins in their habitat to 'protect' them); and the artificially engineered life-form 'Jiggles', who consumed the zoo's fruit and nearly destroyed the zoo when it grew to monstrous size. In Madagascar 3: Europe's Most Wanted, an invention of his called a "nucular" reactor (sic) is used.

As a result of his inventions, he comes off as unhinged, especially when he becomes overly defensive about what he does. Outright called a mad scientist by Skipper, Kowalski often creates things in order to "show them" and has, on few occasions, hidden his devices. In "I Know Why The Caged Bird Goes Insane", he becomes increasingly derailed as the episode goes on after being temporarily bound to a wheelchair, mostly because the nursery he was being held in was right next door to the Science Expo, and he makes a few attempts to break out.

Kowalski can play a banjo and sing as seen in the episode "Concrete Jungle Survival".

Private

Voiced by:

• James Patrick Stuart - in The Penguins of Madagascar
• Christopher Knights - in the movies and other media

Private is the youngest and shortest of the penguins. He has light blue eyes and speaks with a British accent (though the other penguins think his accent is fake). Private has been known to be a rather curious penguin and is much nicer and down-to-earth than the others. In a way, he's much more childish, enjoying "Lunacorns" (pony dolls based on a show that teaches good morals and friendship) and often suggesting much nicer plans (such as asking for something instead of breaking into a habitat or simply telling the truth to others.) He is often called inexperienced by the others. He has an addiction to Peanut Butter Winkies, first displayed in "Skorka" and further elaborated on in later episodes.

He was overprotective of the egg in the episode "Paternal Egg-Stinct", though given how his team mates were treating it (Skipper put it through a dangerous obstacle course, Kowalski tried shock treatments to increase its intelligence, and Rico was flying it with a hang glider), this was rational. Private is very good at deciphering code, communications and peace treaties, and in some degrees, common sense.

Private has been known to give life lessons (in "Mort Unbound" he tells Mort, "There is a natural order of things" and in "Two Feet High and Rising" he tells Mort that Julien's feet "are just feet; not love"). Private is emotionally sensitive, and is often portrayed as more observant or down to earth than the others (noticing when Skipper's ankle tag had not been taken off, indicating that he did not get a shot or suggesting that Maurice had stumbled back because of being blinded by a camera flash, which Skipper quickly labeled as preposterous though he nevertheless told Kowalski to run a scenario) as well as having fighting skills on a par with or only slightly below those of Rico and Skipper (in one episode he is able to block rapid strikes from Skipper and disarm him with his feet while spinning on his head, all while blindfolded). On the wall of the Penguin's HQ is a stuffed fish with a sign "Private's First Prize"; hiding behind it (as well as a bunch of electronic precautions) is his most prized possession: a butterscotch lollipop. Like Kowalski, he loves candy, but eating too much of it causes him to hallucinate. He has an abnormal fear of badgers and cockroaches, as shown in the "Badger Pride" and "Stop Bugging Me" episodes respectively. When Private is the last available penguin to fix a situation (such as when it's revealed that the penguins had spied on the rest of the zoo occupants or during 'Untouchable'), he is able to rise to the occasion and execute a solution that benefits everyone.

In the episode "Cute-Astrophe", it is revealed that Private has a "hyper-cute" ability, wherein he achieves a state of over 132% adorability, that will make everyone around him pass out as they cannot cope with how absolutely cute he is. At first, Skipper abused the hyper-cute, using it on everyone around them, so Private swore to never use it again after it created chaos around the zoo and nearly got them sent away. Despite his vow, the hyper-cute made appearances in other episodes. This ability was attempted in one episode when Private was moulting, but instead of knocking an observer out with adorability, it frightened and panicked anybody seeing him use the ability.

In the "Concrete Jungle Survival" episode, Private received the rank of Private First Class after successfully passing a test in the city. He had been just a Private beforehand.

In the Christmas special "Merry Madagascar" Private finds himself falling in love with Cupid, one of Santa's reindeer. The relationship is short lived, as Cupid must return and work with Santa. Private reminded her that they would "Always have Madagascar!" as she flew off.

Private is the only penguin with known relatives, he has uncle Nigel (though they may not exactly be related), and in "operation penguin" he mentions his father is named Sam Fishy, though Sam Fishy has yet to be shown onscreen, however in Penguins of Madagascar he appeared to have been abandoned as an egg, leading to the possibility that he is an orphan.

Rico

Voiced by:

• Jeffrey Katzenberg - in the first film
• John DiMaggio - in most media
• Fred Tatasciore - in second video game
• Conrad Vernon - in Penguins of Madagascar film

Rico, third-in-command of the group, does not really talk much. He is similar in physical features to the other penguins in the movies, but he has a tuft of feathers shaped like a Mohawk and a scar of unknown origin on his left cheek in the television series. Skipper describes him as a "world-class psychopath". Rico appears to be not merely particularly psychotic, but also mentally deficient. In "Friend in a Box" Kowalski points a mind-reading device at him which, while functioning for everybody else, picks up only the word "Fish" being thought by Rico. According to Skipper, it was a mystery how Rico passed the psych test.

In the first and second film, what little speech Rico uses suggests that his native language is Japanese, and his sushi-preparation skills add to this idea. However, in the TV series, he mostly communicates with various grunts, squeals, and gibberish, (though his language seems to have improved by the second season, as he is able to speak short fragments such as "Allow me" and "You're the King") and has displayed a fondness for a plastic doll toy. He's the team's weapons expert and his main function is to regurgitate any necessary tools that Skipper may need for a mission (as the fact that his stomach operates as a hammer space, several items being as big as himself or weighing a large amount, with no effect on himself or anything/any character he might be on). He's very fond of explosives, and his answer to problems would always be "Kaboom!", if Skipper did not stop him.

Rico has little tolerance for "mushy" stuff, and often becomes sick when Private gets overly emotional and "lovey-dovey" about something. On the other hand he seems to be the second most emotional penguin next to Private and is deeply in love with a doll called Ms. Perky. Another characteristic of Rico is that he might be incredibly superstitious, as, after receiving a rather dismal sounding fortune in a fortune cookie, Rico was a victim of pranks by King Julien who was trying to prove it was right. Even after revealing that Julien was behind the accidents, Rico still needed to have him "take away the curse." He is also insanely skilled with knives and bladed objects, able to make sashimi or carve a Christmas tree from ice in a matter of seconds and destroy bowling pin dummies with a chainsaw while blindfolded.

Rico's psychosis needs to be controlled at all times, by Skipper limiting his use of destructive solutions, often involving explosions. Giving him free rein to destroy to his heart's desire, Rico builds up enough psychotic pressure that he becomes a vortex of destruction, spinning out of control and destroying everything in his path. Kowalski once estimated Rico was capable of destroying the entire zoo in this state.

Rico appears to have outstanding opera voice when excited (as seen in episodes "Go Fish", "Herring Impaired", "High Moltage", "Hair Apparent", etc.).

In episode "Herring Impaired", he is actually most responsible of penguins and because other penguins have "Pisces Dementia", Rico must watch, if no penguin have another fish. That means, he can be serious and responsible when his friends have a big problem.

The Lemurs

King Julien XIII

Voiced by:

• Sacha Baron Cohen - films
• Danny Jacobs - television, specials, video games and "The Penguins of Madagascar"
• Keith Ferguson - first video game

King Julien XIII is a conceited, selfish, fun-loving ring-tailed lemur.

As a self-proclaimed lord of the lemurs, Julien ruled over a great colony of lemurs in Madagascar using mostly his charisma and delegating, i.e. telling other people what they should do since he certainly had very little ability to lead. Nevertheless, he shows more intelligence and calmness compared to the other lemurs (aside from Maurice), which is shown when the very word "Fossa" drives all of them, except himself and Maurice, into a state of panic.

He had parties for his people on Madagascar with much frequency, the likes of which were interrupted constantly by the fossa, which trespassed onto the lemur territory, devouring all that they could catch.

In Julien's early life as seen in All Hail King Julien, he was depicted as the nephew of Uncle King Julien XII, who banned all noisy things to keep the fossa from attacking. After Uncle King Julien XII abdicated to evade a prophecy from being eaten by the Fossa, Julien became the new King of the Lemurs and did things that were opposite his uncle's rules. He saves his people from the Fossa and got bitten on the butt by the Fossa and he survived. When he learns that his nephew is still alive and is still king, his devious uncle tries to get rid of him and reclaim the throne. When he returned, Uncle King Julien tricks Julien into going into the Fossa territory to find out what they're planning with Maurice joining him. In the end, Julien discovered his uncle's plot, glad that Clover chose him over his uncle, and gives his uncle a second chance, by putting him in the Fossa disguise, that is being cuddled by a Fossa girl, much to his uncle's dismay.

The day Alex and his friends came to Madagascar, King Julien and his subjects were enjoying one of their parties when it was invaded by the fossa. When the fossa were scared off by Alex, Julien and his people were hiding in the trees and saw this scene, thinking Alex and his friends were giants from some faraway land, with Julien coming forward and presenting himself as the king of the lemurs â€" once he had ascertained they were not dangerous by tempting them with Mort.

When asked where the people were, Julien pointed to a skeleton of a man caught in a parachute hanging from a tree. Later that night, Julien had a meeting of the lemurs and other creatures of the jungle in the plane. Julien insisted that if they could make friends with Alex and his friends, the foossa would be scared away for good. Maurice was skeptical about this plan, but Julien insisted that, being king, his ideas were the best. Julien's plan failed when Alex started to get out of control and try to kill the lemurs and his friends. Although, Alex then turned back to normal and the foossa were scared away with only the actions of the four main characters and the penguins, Julien took the credits for its success. Interestingly though, when it went wrong, he did not foist the idea back onto Maurice.

When Alex and his friends were ready to go back to New York, Julien gave them his crown; he had already made himself a bigger, better one using a leaf-tailed gecko named Stevie.

In Madagascar: Escape 2 Africa, Julien, along with Maurice and Mort, accompany Alex, Marty, Gloria and Melman to New York on the plane built by the penguins while leaving Stevie in charge until the day he returns. He is in the first class, ordering constant refreshments, and amusing himself with plane crash videos. As the plane crashes down to Africa after running out of fuel and momentum, Julien takes utter pleasure in the feeling of weightlessness caused by the high speed descent. By being in first class, he escapes the crash with a parachute, but loses his crown.

Upon arriving in Africa, Julien at first believes they are in New York, and makes himself a new crown. He manages to gain some power while in Africa, although very few characters actually pay attention to him, getting to ride a flock of flamingos with Maurice, then moving on to riding ostriches and finally an elephant. Later, when the reserve's watering hole dries up, Julien suggests to the animals that they make a sacrifice to "his good friends, the water gods" at the volcano to replenish the water. To convince the masses, King Julien enacts his casual conversation with the Water Gods. Later, Julien persuades Melman to volunteer for the sacrifice, who believed he was going to die soon anyway. But when Melman realized he was going to live, he gave up on being a sacrifice, leaving Julien disappointed at his plan's failure. Mort arrives soon after, followed by a persistent shark that had been attempting to eat him. This shark then falls into the volcano, and as Julien comes out of the volcano, he sees the water coming back to the reserve (due, in fact, to a dam constructed by Nana being destroyed by Alex and Zuba), and is convinced of his plan's success. From this King Julien concludes that the Water Gods like sea food more.

Julien appears in The Penguins of Madagascar regularly, often noisily and unintentionally annoying Skipper and pretty much everyone. He is the self-proclaimed "King of the Zoo" no doubt due to his former king status on Madagascar. Only Mort and Maurice truly believe and respect him as their king, the rest just show up to his "proclamation of royal decrees." Julien is proven to be a worthy ally in most episodes, since it is he who sometimes saves the day for the penguins (like in "Dr. Blowhole's Revenge"). Julien hates having anyone touch his feet, such as passing a royal decree that anyone who touches his feet would be banished from his kingdom (the zoo). When the sewer rats went to the zoo and tricked the Penguins out of their home, Julien decided to play for the Penguins in a game of Hockey to try and win their home back. They were losing until one of the rats touched his feet, after that he lost it and beat all of them on his own.

In the episode "Crown Fools", Julien panics when he loses his crown and obsessively longs for it until it is revealed that he had a spare crown all along (prompting Marlene, who was trying to make him new crowns all day, to chase him with a crowbar). Despite his desire for everyone to listen and do what he wants, he dislikes and ignores Mort, his most loyal follower, who treats him like a god. Mort responds to everything Julien tells him to do as "I like..." followed by whatever Julien said. In the episode "Haunted Habitat" he mentioned that he had a dream that he was "The last mammal on Earth". Mort was just happy to be in the dream despite that he was "road kill" in it.

A recurring gag in The Penguins of Madagascar series is that Julien believes in supernatural beings, which sometimes annoys the other animals. In the episode Misfortune Cookie, Rico gets a fortune cookie that reads "You will soon meet a fowl end." The penguins deny that fortune cookies have any power, despite receiving three accurate fortunes, and dismiss it as superstition. Julien exclaims "these stitions are super," and tries to get the penguins to believe in the supernatural. King Julien actively plots to make a "foul end" fortune come true - by his own doing. It is also known that his believing in spirits can be used against him, seeing that Mason and Phil tricks Julien into believing that the "sky spirits" are angry with the Julien self-centrism, thing that change Julien's personality. But when Skipper notices that kind Julien is more annoying than the narcissist Julien, Skipper makes him come back to his original self.

It is also shown that Julien gains a super-lemur level of strength if he gets mad. Visible when in "Miracle on Ice", Julien spanked and humiliated the Rat King, after one of his henchmen touched Julien's feet; In Happy King Julien Day, Maurice claims that Julien violently used a cane to spank everyone who forgot his self-proclaimed holiday.

In Madagascar 3: Europe's Most Wanted, he has a bigger role and falls in love with Sonya the Bear. Also, he saved the day by managing to get back to the circus.

Maurice

Voiced by:

• Cedric the Entertainer - movies and specials
• John Cothran, Jr. - first video game
• Fred Tatasciore - second and third video games
• Kevin Michael Richardson - in television

Maurice is a "big boned" (overweight) Aye Aye.

Maurice is King Julien's advisor - not that Julien ever listens to him - and is highly skeptical of his ruler's mad plans. He has a long squirrel-like bushy tail. He lived in a tribe of other lemurs and witnessed newcomers to their home: a quartet of zoo animals. Maurice was suspicious of Alex, a lion, thinking him to be potentially dangerous. Maurice expressed this concern during a meeting of the lemurs and jungle animals, but was ignored by Julien. The next day, Maurice took part in the festival honoring the newcomers, and shortly after explained the trouble with having a lion around. He knew that Alex would need to eat meat, and felt that Alex should leave and go to the carnivore side of the island.

Shortly after, Maurice witnessed Gloria, Melman, and Marty working with a group of penguins to fight off the fossa that were attacking the lemur tribe. Maurice had to admit that King Julien's plan had worked and that fossa would no longer be of any worry. Maurice saw the newcomers out when they decided to leave and was obligated to wave King Julien's arm for him, as the king was tired of waving it.

In the sequel Madagascar: Escape 2 Africa, Maurice does not have a prominent role, basically agreeing to whatever King Julien says. In the series, he often acts like a strict big brother to Mort and knows him so well that in "The Penguin Stays in the Picture" he is shocked to find that Mort only ate half of his popsicle, prompting Skipper to go find him because he suspected Mort went missing, which was correct.

He is shown to have many hidden talents in several episodes such as painting and speaking chameleon in "The Hidden" and using a yo-yo in "Friend-in-a-Box." In "The Lost Treasure of the Golden Squirrel," his desire to control an army of Julien manservants is revealed. Despite being overweight, he was still able to hold his own when he got into a physical fight with Kowalski and Rico in "The Lost Treasure of the Golden Squirrel." He has shown attraction to Darla the Baboon.

Maurice returns in Madagascar 3: Europe's Most Wanted.

Mort

Voiced by:

• Andy Richter - movies, specials, The Penguins of Madagascar
• Dee Bradley Baker - first game
• Matt Nolan - second and third games

Mort is a small cute and cuddly goodman's mouse lemur (in the French version, he is a Bushbaby).

Though quite timid, Mort can also be shrill and vocal in confrontations. Julien finds Mort incredibly annoying, and does not hesitate in proclaiming so. Despite this, Mort does not show any lasting unhappiness, and his expression changes within the same scene. A running gag is that Mort commonly pukes. Mort has an obsession with Julien's feet ever since he used them to kick some fossa as seen in the first episode of All Hail King Julien. Mort also has very good money skills as in the episode "Gimme Gimme Gimme: The Game" where he cleverly becomes the richest lemur in Madagascar (all the other lemurs became broke including Julien after getting all their money and possessions but gives it all back after Julien, defeated, says that he had won showing that he believed that it was all a game).

In The Penguins of Madagascar episode "Sting Operation" when the penguins turn into idiots, they had the same obsession for feet just like him, which Julien despises. In his anger, Julien once banished Mort from their habitat at the zoo, forcing Mort to seek a "cure" from the Penguins.

Mort makes his first appearance in Madagascar, wherein he lives among the other native lemurs under Julien. Julien uses him as a test to see if Alex and the others eat lemurs, and Mort quickly decides that he likes them, as he is later shown to have a tendency to like everything. Later, Julien notes that if Alex does not like fish and still hungers for steak, Mort is "Plan B". Mort is last seen in the film with the other lemurs wishing the zoo animals farewell, and Julien uses him as a makeshift foot-rest.

In Madagascar: Escape 2 Africa, Mort plays a much smaller role. He attempts to follow Julien and Maurice on the penguins' plane, but Julien goes to extremes to keep him out of the plane, declaring that Mort had "scissors and hand cream" (he did in fact, have a pair of scissors). Mort succeeds in attaching himself to the outside of the plane, and promptly falls into the ocean shortly before it crashed. After reaching a shore at Africa, Mort is pursued by a persistent shark, which follows him onto the African preserve. Mort (still followed by the shark) finds Julien at a volcano where Melman had just decided that he would not sacrifice himself to return the preserve's water. The shark accidentally falls into the volcano just as Alex and Zuba destroy the dam blocking the river, thus leading Julien to believe that his plan to give a sacrifice to bring back the water was successful.

Mort makes appearances through The Penguins of Madagascar, along with Julien and Maurice. At one point, stuffed "Mort" dolls became very popular, and out of jealousy Julien throws them into the skunk habitat. Mort is accidentally shipped back to the factory with the dolls, and must be rescued. In the episode, King Julien also reveals that the only reason he allows Mort to stay with them is so he can express his dislike for Mort in front of him, and keep himself entertained. Mort is also shown to be unhappy with his past self after growing in size and muscle mass due to an invention from Kowalski. After being turned back into his regular self, Mort expresses that he is happy with who he is. In the television series, whenever an animal or object is falling it almost always lands on Mort. Despite the fact that Mort is often treated poorly by King Julien, the other characters show concern for him as Maurice tries to (if with limited success) save Mort when he may be in danger to be King Julien's biggest concern, and Skipper, who refers to Mort as "sad-eyes", dived in the way of the mutant rat overlord to save Mort. In the episode "Otter Woman", all the boys are attracted to Marlene because her fur turned white. Mort tries the same thing in the end of the episode, without success.

Mort returned in Madagascar 3: Europe's Most Wanted. Things he does in it involve vomiting on Alex's birthday cake, helping Alex defeat Dubois early on in the film, being part of the flying Afro Circus with the other lemurs, and tranquilizing Dubois with knockout darts.

Mason and Phil

Voiced by:

• Conrad Vernon - Mason (Phil has no dialogue)

Mason and Phil are two sophisticated common chimpanzees at the Central Park Zoo, who prefer to drink cups of coffee and "read" the newspaper in the morning. Mason speaks with a stereotypical British accent, while Phil doesn't speak and communicates through American Sign Language. While Mason is unable to read, Phil can and is used to decipher writing while Mason translates the ASL. Both escape the zoo after Gloria breaks through its outer wall; Mason mentions Tom Wolfe giving a lecture, and they plan to attend to "throw poo at him." Along with other characters, Phil and Mason are captured and sent to a wildlife reserve. However, their crates are not thrown off the boat, and they are not seen until a final scene, in which they are still in crates waving farewell to the Madagascar natives.

In Madagascar: Escape 2 Africa, the chimps call upon chimpanzees living in Africa to help rebuild the wrecked plane that was supposed to fly them back to New York. They later go on strike, asking for maternity leaves and breaks (Skipper argues that they are all males); eventually, they blackmail Skipper into agreeing.

In The Penguins of Madagascar, Mason and Phil are supporting characters in the series, and are shown to be annoyed by Julien's partying. They attempt to rid him of his ways by tricking Julien into thinking a solar eclipse was a sign that the "sky spirits" disapproved of his behavior. They also frequently mention throwing poo and are used to read the English language. Phil may be the most temperamental or "potty mouthed" of the two, as after being told there is no checkmate in checkers, he makes several signs which Mason responds with "You groom your mother with those hands?!" Phil and Mason may also sometimes lose things in the translation of what they are made to read. An instance of this is when the zoo occupants were ordered to make a cake for King Julien, and Mason mistakes "booger" for "sugar". This does not mean that Phil is unable to communicate well, even though he is unable to truly speak, he was still able to win over the heart of Lulu, a female chimp visiting from the Hoboken Zoo in New Jersey for a weekend, While Lulu did appear in a later episode where the penguins ended up accidentally finding themselves in the Hoboken Zoo, it is unknown if she keeps in touch with Phil.

McGrath explained that originally their "first scene was just deciphering the code on the shipping label". The crew "got this girl who knew American Sign Language" to give them hand gestures. They wanted gestures to be "really frenetic", and "she signed out, 'Tell the tiny pea-brained birds that the sign reads: Ship to...'". Afterward, the crew returned to the scene where Mason notes Tom Wolfe's lecture, and she signed "Can we throw our poo at his stupid white suit?" Mason is named after legendary British actor James Mason, whose urbane, sophisticated vocal styling is mimicked by Vernon.

In Madagascar 3: Europe's Most Wanted Mason and Phil have smaller roles and are mostly seen posing as the King of Versailles.

They make a cameo appearance on a newspaper in Turbo.

Circus Zaragoza Animals

The animals of Circus Zaragoza are trained circus performers that Alex's group had met while on the run from Chantal DuBois. Among the animals of Circus Zaragoza are:

Vitaly

Vitaly (voiced by Bryan Cranston) is a Siberian tiger with a Russian accent who leads a traveling circus in Madagascar 3: Europe's Most Wanted.

The tough Vitaly once jumped through flaming rings of fire to excite crowds, constantly pushing himself to the limit by jumping through smaller and smaller rings that seemed impossible for his size while lubricating himself with olive oil. But after losing his confidence and more than a few of his hairs on one fateful attempt which involved a flaming ring that would only fit on the smallest finger (due to the olive oil being flammable), he also lost his passion and his act partially. When Alex and his friends plead with the circus to let them escape on their train, Vitaly is the most resistant, only to be overruled by Gia. When the zoo animals purchase the circus, Vitaly remains the most hostile to their changes to the business, even as the rest of the crew becomes inspired.

In London, Vitaly decides to give up and prepares to leave just before the circus' critical performance. Alex confronts Vitaly and reminds the tiger of his love of performing and while suggesting he uses hair conditioner as a safer lubricant before regaining his courage to jump through the tiny flaming hoop. As a result, Vitaly's stunt is performed perfectly to open the successful show. With that, a profoundly heartened Vitaly becomes a fast friend of Alex. Even after the zoo animals are exposed and estranged from the circus, Vitaly is the first to support Gia's proposal to rescue their new friends from the zoo and plays a key role in that operation.

Gia

Gia (voiced by Jessica Chastain) is a jaguar with an Italian accent who is part of the traveling circus in Madagascar 3: Europe's Most Wanted. She wears a blue woven floral necklace, has gold eyes and has heart-shaped palms. She is Alex's love interest who is always on the lookout to discover something new and gets her chance when she convinces a reluctant Alex to teach her the trapeze â€" "Circus Americano" style. Her only role in the circus before Alex, Marty, Melman and Gloria came around was doing normal cat tricks (sitting, standing, rolling over), but when Alex teaches her trapeze, she performs with him and they move closer. Later on, she and the other circus animals save Alex and the gang from Dubois.

Stefano

Stefano (voiced by Martin Short) is a New Zealand sea lion with an Italian accent and curly whiskers. He is part of the traveling circus in Madagascar 3: Europe's Most Wanted and is a happy, fun-loving Italian sea lion who always looks on the brighter side of things. A true entertainer at heart, he is up for any challenge to save the circus, even convincing Alex that the show must go on.

Sonya

Sonya (vocal effects provided by Frank Welker) is the traveling circus’s most ferocious Eurasian brown bear who appears in Madagascar 3: Europe's Most Wanted. Her tricycle riding act is only topped by her ridiculous tutu. But her biggest trick of all will be casting a spell on King Julien, who is instantly smitten with her. She does not talk, but Julien can understand what she is saying when she growls (the way she communicates instead).

While they are in Rome, she and Julien travel to Vatican City where they steal the Pope's ring after Julien pretends to kiss it and uses it to buy her a Ducati motorbike after they fall down a set of stairs and destroy her tricycle, which she uses in her performance in London.

The Andalusian Triplets

Esperanza, Esmeralda, and Ernestina (voiced by Paz Vega) are three of the new characters in Madagascar 3: Europe's Most Wanted. They are white Andalusian Mares who like Marty and are part of the circus. All their names start with "E" and end with "A". When Alex's group improved the Circus Zaragoza, the Andalusian Triplets bounced on the trampolines during their act while wearing butterfly wings on their outfits.

They were listed as "Horses" in the credits.

The Dancing Dogs

The Dancing Dogs are a group of six performing dogs with British accents as members of the circus in Madagascar 3: Europe's Most Wanted. They tend to fight and argue with each other and they hate being called cute and cuddly. The Dancing Dogs consist of Frankie the Entlebucher Mountain Dog (voiced by Nick Fletcher), Freddie the Great Dane (voiced by Vinnie Jones), Jonesy the Bernese Mountain Dog (voiced by Steve Jones), Shakey the Brown Bernese Mountain Dog, and two unnamed dogs. Only the first three are credited.

Manu and Maya

Manu and Maya are two performing Indian elephants as members of Circus Zaragoza in Madagascar 3: Europe's Most Wanted.

Their names are revealed in the DVD commentary as well as the fact that they are German. According to Tom McGrath, they are very nice and great to work with. They remain silent in the film due to the fact that their possible dialogue was cut from the film.

Secondary characters

Ted

Ted (voiced by Bill Fagerbakke) is a polar bear who lives in the Central Park Zoo. He first appeared in "The Madagascar Penguins in a Christmas Caper" where he was sad due to be alone during Christmas. This caused Private to want to give him a present. When Skipper denied him his request, Private snuck out into to the city to find Ted a gift only to be kidnapped by Nana. After getting Private back, the other penguins decided to invite Ted to their Christmas party where the polar bear had invited the rest of the Zoo as the animals present sing their rendition of "Jingle Bells."

He made a brief background cameo in the first movie.

In the series, he mainly appears as a background zoo animal. He did play a small role in the episode "P.E.L.T." when Skipper had to proke and evade a polar predator.

Zuba

Zuba (voiced by Bernie Mac in the film, Dan White in the video game) is a lion who is Alex's father and the alpha lion of the pride in Madagascar: Escape 2 Africa.

He was first seen as the Alpha Lion of a pride in the reserve where Alex was born. At first he thinks that Alex has returned to take over, but his wife notices that they both have the same birth mark and realizes that Alex is their son. Zuba is a proud father, but he is somewhat confused as to why his son is interested in dancing. After many events in the movie including Alex failing the rite of passage and shaming his family, Zuba finally accepts Alex. The film was dedicated to Mac's memory.

Florrie

Florrie (voiced by Sherri Shepherd) is Alex's mother, Zuba's mate, and the alpha lioness of the pride in Madagascar: Escape 2 Africa. Florrie, unlike Zuba, accepts Alex for what he is. She is not disturbed at all by the fact that Alex prefers to dance instead of fight. Due to this, she acts as the voice of reason and tries to encourage Zuba that it is better that Alex came back to them, dancer or not, as a king. Florrie is never addressed by her name in the film and in the movie's end credits, she is credited as "Mom". It is presumably in the novelization that her name is exposed.

Moto Moto

Moto Moto (voiced by will.i.am in the film, Greg Eagles in the video game) is a big-bellied, handsome, muscular hippopotamus with a deep voice, whose main hobby is flirting with anything of the female species. Moto Moto is kind, big, chunky and charming. He is described by him as being "aggressive but friendly", and is the way the standard male hippo should look. In Madagascar: Escape 2 Africa, he falls for Gloria as a major plot boost. When they're dancing in a lake, Gloria realizes that Moto Moto only loves her because of her large body. Gloria breaks up with him as a result, though they remain friends.

Moto Moto means "Hot Hot" in Swahili.

Okapi Girl

An unnamed female Okapi (voiced by Taraji P. Henson) appears in Madly Madagascar, whom Marty finds himself in love with. She is not interested until he uses a love potion on himself, causing her and all the other female animals to begin pampering him. She and the other animals lose interest in him after he jumps into a lake to hide from them (due to becoming overwhelmed by the pampering), causing the potion to wash off.

North Wind

North Wind is an Arctic elite undercover interspecies task force that is dedicated to helping animals that can't help themselves. They worked with the Penguins in order to thwart the plans of the villain Dave (aka Dr. Octavius Brine). Among the members of North Wind are:

Classified

Classified (voiced by Benedict Cumberbatch) is a courageous yet slightly arrogant British gray wolf and the team leader of North Wind. He is at loggerheads with Skipper. Skipper calls him Classified, who states his name isn't really "Classified". His name is Classified because he is the leader of the team. He is the North Wind's counterpart of Skipper.

Short Fuse

Short Fuse (voiced by Ken Jeong) is a Belgian harp seal. He is the North Wind's explosive and demolitions expert.

Eva

Eva (voiced by Annet Mahendru) is a Russian snowy owl and North Wind's intelligence analyst. Kowalski has a crush on her.

Corporal

Corporal (voiced by Peter Stormare) is a Norwegian polar bear and the muscle of the North Wind team. He seems to like penguins and he freaks out when other penguins are taken.

Antagonists

Nana

Nana (voiced by Elisa Gabrielli in the films, Marion Ross in the second video game) is an elderly lady with a German accent and the archenemy of Alex the lion.

Nana's personality changes over her appearances. Initially, she appears as a very aggressive old lady with incredible martial arts skills. In The Madagascar Penguins in a Christmas Caper, she is shown to be aggressive with anyone, no matter how courteous people are with her. But in Madagascar: Escape 2 Africa, she appears to be kinder than before, even leading the New Yorkers when she notices that they are in panic, only being aggressive with wild animals; which she sees as no different from domestic animals. She is also shown to be a soft and grandmotherly figure to the New Yorkers. For example, when a man helps her find her handbag, she says, "Such a good boy. Nana can't survive without it."

Nana seems to be incredibly resistant to injury, as she was able to have another fight with Alex right after being tugged out of the back of a speeding tour jeep as well as being thrown through the windshield a second time by the penguins, who reversed over her in the jeep after noticing that she did not die. Despite her martial arts skills, she is still portrayed as a "little old lady"; she has a waddling gait, wears dentures, and is knocked onto her back when firing a gun.

In the first movie, Nana is a minor antagonist when she crosses paths with Alex in Grand Central Station, then brutally assaults him. Throughout the series, she refers to Alex as "the bad kitty." She also hits Melman with her hand bag, causing his head to get stuck in the Grand Central Station Clock. When the animals are surrounded by police, she somehow gets past and kicks Alex in "the batteries" (or his groin). She was quickly taken away by police. As shown in the sequel, she was not arrested, she was interviewed as a witness, and states what she thought of Alex: "He was a very bad kitty."

Nana is featured in The Madagascar Penguins in a Christmas Caper, where she proves that there are times when she is shown to be not only aggressive with animals, but with also people, like when she destroys random merchandise at a kiosk while shopping for her dog, then demands to purchase Private in a very rude tone, when she refuses to pay the taxi driver her fare, when she assaults her doorman for no reason, and even when she does not hold the elevator for Skipper, Kowalski, and Rico.

Nana returns in Madagascar: Escape 2 Africa, as the secondary antagonist. She also becomes the leader of an army of tourists left stranded by the penguins' schemes. She again crosses paths with Alex, then fights him, and wins again, although Alex puts up much more of a fight this time (although Alex confesses to his friends that he actually only did it to distract her to get her bag, making Nana's victory unofficial). She often hits Alex and loves her handbag. At the end of the film, Alex gives Makunga Nana's handbag. When she sees Makunga holding it, she kicks him in the groin, steps on his foot, whacks his hand with a ruler, digs her hand into his ear, arm-burns him, and spanks him before dragging him off the reserve by his ear. According to the DVD commentary, she took Makunga back to New York with her in a kitty cage.

Nana's assault on Alex at Grand Central Station is mentioned by a taxi driver in The Penguins of Madagascar episode "Zoo Tube".

Nana does not appear in Madagascar 3: Europe's Most Wanted.

Mr. Chew

Mr. Chew is a small, white poodle, fond of chewing and tearing various items apart (as demonstrated on an Alex plush toy). Chew is owned by Nana, and lives in her apartment on the top floor in an apartment complex. Mr. Chew is the main antagonist in "The Madagascar Penguins in a Christmas Caper". Nana purchased Private, mistaking him for a squeaky toy, while he was attempting to find a gift for Ted, and reveals him to be a Christmas present for Mr. Chew. Chew nearly mauled Private, while Nana was watching a football game on TV, but was defeated by Skipper, Kowalski and Rico, who had set out to rescue Private. Nana does not hear the fight, but after she sees the damage (following hearing Rico's dynamite stick going off), she blames Mr. Chew for destroying her apartment and puts him on a big "time out".

Mr. Chew had a cameo appearance in Madagascar 3: Europe's Most Wanted when an identical poodle was seen as a head trophy owned by Captain Chantel DuBois.

The Fossa

The fossa (voiced by Tom McGrath and Eric Darnell in the film, various voices in All Hail King Julien), are the main antagonists of the first film and their name is misspelled "foosa" in the film. They appear as medium-sized weasel-like creatures, closely related to mongoose, and rarely speak, instead usually growling and yelping.

Throughout Madagascar, they are shown to terrorize the lemurs and attempt to eat them. According to Julien, "they're always annoying us [the lemurs] by trespassing, interrupting our parties, and ripping our limbs off." The fossa are fearful of Alex, and run away when they suspect an attack from him, as well as allow him to take their prey. The fossa are defeated by Alex, Marty, Gloria, Melman, and the penguins and driven into their territory.

In "The Penguins of Madagascar" episode "The Terror of Madagascar," a baby Fossa (voiced by Dee Bradley Baker) was shipped to the Central Park Zoo as part of a zoo-to-zoo tour. Throughout the episode, the baby fossa kept biting King Julien. Because of this, he thought that the fossa was trying to eat him. Kowalski told him that this was the fossa's way of showing love towards Julien, since he though of him as his father. But, Julien realized this too late, because in order to get rid of the fossa, he shipped in a boa constrictor named Savio to devour him. In order to keep the fossa safe, Julien wanted Savio to eat him instead, but the fossa attacked Savio to protect Julien, much to the penguins' disappointment.

The Fossa appear in "All Hail King Julien" as recurring antagonists. During his reign, King Julien's uncle King Julien XII had kept the lemur kingdom quiet in order to keep the Fossa from attacking them. His uncle gives Julien the crown after learning that the king of the lemurs will be eaten by the Foosa. But Julien got bitten on the butt, while saving the lemurs and his scream sent boulders falling down and the Foosa run away. Throughout the first movie and the two TV series, there have been male Foosa. In the fifth episode of All Hail King Julien there is a Foosa girl who flirts with Julien and Maurice in a Foosa suit. She is seen in the end cuddling the suit with Julien's uncle in it who is being punished and sparred by Julien, for sending his nephew to the territory and trying to reclaim the throne.

Makunga

Makunga (voiced by Alec Baldwin in the film, John Cygan in the video game) is a male lion with a large black mane styled in a pompadour (similar to Elvis Presley's hair) and green eyes, and is the main antagonist of the second film. He is Zuba's rival and wants to take his place as alpha lion. He also calls Alex pathetic. When asked why he wants to become alpha lion, he replies "I'm better looking, I have better hair, I'm deceivingly smart, and I want everyone else to do what I say" Makunga usually acts like he thinks he is smart and has good leadership, but he only is trying to become alpha lion because he wants to be in charge. Makunga often looks at other characters while making an evil unnoticed face where he crosses his arms in front of his chest and makes a big smirk , usually when he is planning to do something that is not good. (he is very similar to Scar from The Lion King")

His name sounds a little to "Wakunga", which means "midwife" in Swahili.

Makunga often challenged Zuba to fights over the alpha lion position, but was easily beaten. One of those fights led to Zuba being distracted while Alex was captured by poachers. Years later, Makunga used Alex in a bid to finally reach the rank of alpha lion where he tricked Alex into challenging his henchman Teetsi. When Alex was defeated, Zuba reluctantly had to give up his alpha lion status and go into exile with his family. Makunga is not a very good leader saying the only solution to the fact that the local water hole was nearly dry was that they all would have to fight for it. This causes doubt about his role as alpha lion, and almost every animal on the reserve wants Zuba back. After the water was restored, he still refused to allow Zuba and Alex to return from banishment. He eventually meets his downfall when being tricked into angering Nana, who attacks Makunga and drags him off the reserve by his ear. The DVD commentary reveals that "Nana brought Makunga to New York in a kitty cage".

Teetsi

Teetsi (voiced by Fred Tatasciore) is Makunga's henchman and the strongest lion in his group. Makunga tricked Alex by suggesting that he goes up against Teetsi. Alex ended up defeated by Teetsi. He is an antagonist in the second film.

Captain Chantel DuBois

Captain Chantel DuBois (voiced by Frances McDormand) is a Monegasque animal control officer and the main antagonist of the third film. She is the best animal control officer in Monte Carlo with a perfect success record. She is a big-game hunter and has extraordinary tracking skills, as well as superhuman strength and a habit of mounting the heads of every animal she has caught on her wall. Like Nana, she is incredibly resistant to injury. Throughout the third film, she pursues the Four in hopes of having their heads as trophies, especially Alex's.

She finally captured Alex and the other animals at New York City, and the zoo staff thank her incorrectly believing that she tried to return Alex. She rips up the check they give her where she says that "It wasn't about the money. It was about the lion." Alex is caged and DuBois hides a poison filled dart in a foam finger and shoots it at him, but the circus animals, being alerted by King Julien, arrive and rescue him. After a long fight between the animals and DuBois and her henchmen, DuBois is tranquilized by Mort and her men are knocked out. She and her men are last seen shipped to Madagascar with her henchmen (not similar to the first movie) which the Penguins did to them.

DuBois' Men

DuBois' Men are four men of Captain DuBois animal control officers, who are antagonists of the third film. They are prone to getting hurt, unlike their boss. DuBois heals them instantly simply by singing "Non, je ne regrette rien" and appealing to their patriotism. One is short, one is tall and skinny with a mustache (on an extra released on the DVD reveals his name is Gerard), one is overweight with a goatee and sideburns, and one is also overweight and has a mustache.

Dave/Dr. Octavius Brine

Dave (voiced by John Malkovich) is a villainous and disgruntled octopus who serves as the main antagonist of Penguins of Madagascar. He disguises himself as a human professor under the alias of Dr. Octavius Brine. Dave used to be one of the top attractions at the Central Park Zoo, in which time he entertained children with his clever and funny tricks. However, when Skipper, Kowalski, Rico and Private arrived, they unknowingly stole all his glory due to their cuteness and he was forcibly removed from his home. Afterwards, Dave was shipped to every major Zoo in the world, but the same thing happened again and again and again: penguins getting all the attention while Dave was shunned and ignored. Embittered, Dave now seeks revenge on the penguin species, becoming a mad scientist in the process.

Inside his submarine lair, Dave creates the Medusa serum, a green substance that when fired in a ray will transform its target into a physically and mentally disfigured monster. He plans to use it on all the penguins that stole his fame and then unleash the now monstrous penguins on New York city, so that the humans will despise them instead of love them. However, a group of Arctic animals called the "North Wind" attempt to foil his plans.

At the end of the film, Dave's plan for revenge is foiled and the vindictive octopus is shrunk by his own ray (the effects having been reversed by Private) and gets trapped in a snow globe, where he is admired by a little girl.

Dave's Henchmen

Dave's Henchmen are octopi who look like squids. They are antagonists of Penguins of Madagascar. They are evil minions who work for Dave and assist him in his plans to rid of all penguins. They don't speak, but they make a bubbling sound that Dave understands. Each henchman share the first names of famous celebrities. Whenever Dave ordered them to do something, he would say the actor's first name and an action word that spells out the actors' full names (Ex. "Nicolas, cage them." "Elijah, would you please take them away?" "Halle, bury them."). About three of these underlings are named after the actors.

Characters in The Penguins of Madagascar TV series

Marlene

Marlene (voiced by Nicole Sullivan in normal form, Dee Bradley Baker in wild form) is the Central Park Zoo's only resident Oriental small-clawed otter with primarily brown fur, with a white chest, face, and right foot. Marlene is shown to be both playful and kind towards others, which often conflicts with Skipper's serious and safety-cautious attitude. Though this is true, the two still share a very strong friendship with one another and still have the ability to keep that certain bond exceedingly strong, making it rather difficult to break. She attempts to find and bring out the best in others through kindness, though is not immune to feeling anger towards Rhonda, a rude walrus apathetic to others, and works to have her transferred (she regrets the decision after learning Rhonda would be sent to a polar bear reserve, and works doggedly to send her elsewhere). Despite having doubts on the penguins' ways of dealing with issues, Marlene still believes that they can accomplish very difficult tasks. Marlene is known to snore very loudly while asleep, which invokes fear in an alligator named Roger residing in the sewer system beneath her enclosure. Like other residents at the zoo, Marlene is shown to love both popcorn and candy. In order to get candy, she is bribed by Maurice to celebrate King Julien Day and participate in several tasks to keep Julien content. Marlene has also revealed that she won the Internet popularity contest at her past zoo three years in a row (though was unable to compete at the Central Park Zoo due to a scandal) and that she considers Christmas in July "the biggest holiday of the year".

In "Otter Gone Wild", it is revealed that she was born in captivity. When she sets foot in the outside world, she becomes wild. In her primal state, she briefly becomes infatuated with King Julien, and is capable of posing a significant threat even to the equally-psychotic Rico in a fight, later successfully defeating two enraged badgers who had just given the entire penguin team significant trouble.

In "Otter Things Have Happened", the penguins invented a machine that is able to locate anyone's ideal match at a moment's notice; the birds misread the signal her saliva sample gave and present her with Fred the squirrel, upsetting Julien who kidnaps the little guy (who Marlene has actually started to like). When she realizes Fred is not good for her, she dumps Fred and goes back to her single lifestyle. Kowalski, upset that his invention is faulty, throws it out. In the trash the machine begins to show the results of Marlene's test again; we see Fred in the area it was picking up, but then are treated to the introduction of a brand new character, Antonio or the "Otter Bachelor of Central Park", a buff, Latino male otter who has a fancy to frequently playing his Spanish guitar. It is not finalized, but it seems that Antonio was the individual who Marlene's test picked up, not Fred. It is not known yet if the penguins know of Antonio's existence or if Marlene will get to meet him in future episodes but in "The Lost Treasure of the Golden Squirrel" it is shown that Marlene dreams of being in a spacious room surrounded by many otters very similar (if not exactly like) Antonio all playing Spanish Guitars.

In "Dr. Blowhole's Revenge", it is shown that she believes Dr. Blowhole is not real. In "The Return of the Revenge of Dr. Blowhole," she saw that he was real, though made no comment concerning any prior disbelief.

During "The Otter Woman", an over-chlorinated pool resulted in Marlene's fur being bleached white, causing her to be mistaken for an arctic mink called "Arlene," resulting in Skipper and Julien becoming infatuated with her before Skipper became convinced that "Arlene" had done something horrible to the seemingly missing Marlene. Later, Marlene's fur was dyed back to its original color by Alice when she noticed what happened to Marlene.

In "Littlefoot," Marlene was separated from her feral self thanks to Kowalski's latest invention, but the separation resulted in her monstrous alter-ego going on the rampage while 'regular' Marlene was now excessively paranoid and afraid of everything around her. Despite Officer X capturing Marlene's feral self and taking her to animal control after feral Marlene went on a rampage that destroyed most of New York- earning the name 'Littlefoot' in the process-, the penguins managed to recombine Marlene's two selves, making Officer X lose his job again, and the experience of confronting her other self allowing Marlene to gain greater control over her feral side.

In "Badger Pride," Marlene had to put up with the attacks from Becky and Stacy Badger (who are prone to having to put up with badger stereotypes) and her wild side comes in handy when it came to subduing them.

Alice

Alice (voiced by Mary Scheer) is a zookeeper exclusive to The Penguins of Madagascar. She seems to dislike aspects of her job, and is happy to have a voice-automated robot guide give directions so she can avoid "annoying tour group questions". She also does not understand what people enjoy about watching animals on their computer, saying that they do not do anything interesting. Alice is involved in several aspects of the animal's well-being. It's hinted that deep down Alice cares for the zoo animals and only wants what's best for them as she hates it when people feed the animals popcorn, and posts several signs throughout the zoo after catching a child feeding Marlene a kernel. Though this could just be herself keeping her job safe. Alice is also in charge of gathering the penguins for visits to the veterinarian, and later remarks that they have a dentist appointment (Private then confusedly states that they do not have teeth). Alice and the penguins have a mutual dislike for each other, and she believes that they are scheming (though she is actually correct in this assumption). Though another zookeeper can be heard over her walkie-talkie, voiced by John DiMaggio, Alice is usually the only one ever seen on-screen but occasionally an unnamed zoo worker appears sporadically, though his face is always hidden. Unintentionally, Alice won the zoo's internet popularity contest when a camera documenting the animals was accidentally pushed toward her dancing and slapping her butt in the kitchen. In "Miss Understanding", she remarked to a know-it-all kid that the zoo had three male penguins and one female, which turned out to be false (not that she knew or cared, either of which could be true).

Pervis McSlade

Pervis McSlade (voiced by Gary Cole) is the zoo commissioner of the Central Park Zoo.

Pervis McSlade first appeared in "In The Line of Doody." He is the commissioner for the Central Park Zoo. He was making an appearance to announce the opening of the Children's Zoo. Skipper saved him from being hit by a pigeon named Frankie.

He later appears in "The Hoboken Surprise" to make Frances Alberta the new zookeeper of Central Park. When he finds out Frances was building bio mechanical androids to replace the animals with, he fired her.

Zoo Doctor

This unnamed doctor (voiced by Brian George in an Asian-Indian accent) works as a veterinarian at the Central Park Zoo. He works in the "animal care" building. Unlike Alice, he seems to be quite fond of animals. He is bald and wears a white lab coat. Skipper often voices dislike of him because of his use of needles and when injured, he has been known to try and downplay the injury rather than visit the vet.

Chuck Charles

Chuck Charles (voiced by Jeff Bennett) is a newsman who works at Channel-1.

In "The All Nighter Before Christmas," he mentions that he is so used to talking on television that he has forgotten how to talk normally.

Due to an unknown reason in "Goodnight and Good Chuck," he was fired and replaced with his rival anchor Pete Peters and he got a new job at the zoo as Alice's assistant. But he was sat on by Burt, swung out of control by Bada and Bing, tried to get a chameleon off of his face, being dragged away by a vehicle that drove itself, almost got flushed down a toilet, jumped on by Mort, and got sprayed by the amnesia spray by the Penguins. He tried to get his old job back by exposing the Penguins' operation to the world and he got past Rico, Private and Kowalski. The disc was taken by Pete Peters and Peters aired it. But when the world thought the Penguins's operation was fake, Pete Peters was fired and Chuck Charles was reinstated.

Rat King

The Rat King (voiced by Diedrich Bader) is a genetically enhanced, muscular lab rat who resides in the sewer. He is a common antagonist of the penguins. He constantly torments the penguins and never learns his lesson when he's defeated each time. He even tries to take over their home in a hockey game, but he is defeated by King Julien after Rat King's fellow rats skated over King Julien's feet. Also he is matched power with Skipper.

Sewer Rats

Several small sewer rats make appearances on the occasion as the servants of Rat King. They are voiced by Jeff Bennett and Kevin Michael Richardson.

Officer X

Officer X (voiced by Cedric Yarbrough) is a New York animal control officer who has a single-minded focus on removing stray animals from the streets. Officer X is shown to have a superhuman level of strength, crushing a small metal cage with his bare hands and defeating Joey, who even the penguins could not easily beat, head on, leading Skipper to say Officer X cannot be human.

He first tangles with the penguins when they interfere with his pursuit of a stray cat, the penguins' friend Max, in the episode "Cat's Cradle". When asked by Alice what his real name was, X responds that his mother never told him his real name, instead telling him that it was "classified." Officer X was defeated by a Red Rhodesian Slasher that was painted to look like Max.

X returns in the episode "What Goes Around" when he tries to capture the penguins when they are found out of the zoo on the streets of New York, but the subsequent chase results in X losing his job due to the scale of the damage he had caused and the public's lack of belief that penguins were responsible for his actions. He was arrested on vandalism charges creating the mess resulting from his and the penguins' encounter.

He returns as an exterminator in "Stop Bugging Me," having been forced to work as an exterminator after his previous rampage. He fights with the penguins who are protecting three cockroaches that Rico befriended. X was ready to enact his revenge, only for Private to cause the cockroaches to crawl into his suit causing him to run off. He also lost his mustache in this episode.

In "The Officer X Factor," he replaces Alice for the weekend when she was sent on a vacation. As a zookeeper, he knew about every inch of the zoo to prevent the penguins' escape into cooler water. He was also expecting that the penguins were going to fly to the East River, but he was defeated there. He landed on the taxi that Alice was returning in as it drives away.

Officer X attempted to regain his position in "Littlefoot" by capturing the monster currently attacking New York (the feral version of Marlene that had been separated from her normal self), but the penguins sabotaged this effort by recombining the two Marlenes, leaving Rico behind to give the impression that X's penguin-related "insanity" had prompted him to steal a penguin from the zoo. X loses his job again, leaving him upset.

His next appearance was in "A Kipper for Skipper" where he was working as a fishmonger. When Kowalski, Rico, and Private were arguing over the fish, X captured the three of them. Luckily, they were able to defeat him by dropping a bowling ball on him.

He was briefly seen in "Goodnight and Good Chuck" in which a video revealing all of the penguins' covert operations were broadcast worldwide. Officer X watched the news segment in Times Square and truimphantly told anyone who would listen that "I told you!" and "I'm not crazy!". He then calls his mother and tells her to turn on the TV to prove his sanity. Unfortunately, everyone else who watches it believes it to be a news hoax due to the sheer impossibility of the scenario and the amusing antics of the penguins in the video. Officer X drops to his knees and cries out in despair when still no one believes him.

Joey

Joey (voiced by James Patrick Stuart) is a bad tempered and hostile red kangaroo with an Australian accent that always refers to himself in the third person who loves beating up anyone who steps foot in his habitat, especially Skipper and Julien. He first appeared in the season one episode, "Assault and Batteries" and later on in "Cat's Cradle" when he tried to beat up Officer X but was badly hurt instead. He also appeared to have a big role in "Skorca!" Though usually only a minor character in the series, Joey has had several episodes where he was the main antagonist. In the season two , episode "Kanga-Management", the Penguins accidentally destroy the Koala habitat, forcing Leonard to temporarily live with Joey until the pen is re-built, to both's dismay. Eventually, the two settle their differences and become friends, agreeing on their hatred for the penguins.

Even though Joey has a pouch that female kangaroos have, it is assumed by fans that Joey is a female kangaroo with a masculine voice.

Leonard

Leonard (voiced by Dana Snyder) is an extremely paranoid and nervous koala. He first appeared in "Night and Dazed", where he was shown to be very afraid of the penguins. After being accidentally launched into the city due to Kowalski's evacuation launcher, the penguins come save him and bring him back to the zoo.

He appears again in "Kanga Management" in which the penguins accidentally destroy his enclosure, forcing him to temporarily live in Joey's enclosure. He and Joey eventually become friends due to their hatred for the penguins.

In "Nighty Night Ninja", he watches a martial arts film with the penguins while he waits for his home to air out, due to a fish bomb. This causes him to start attacking the other animals in a sleep-walking state the next day. The penguins cure him by making him watch The Lunacorns, and save him from the sewer rats, whom he had attacked earlier.

Hans

Hans (voiced by John DiMaggio) is a puffin with a past history with Skipper that involves a mission in Denmark that somehow resulted in Skipper being declared Public Enemy Number One in the country in question. During "Huffin & Puffin", he appeared in New York, initially apparently wanting to make peace with Skipper, before his true agenda was revealed to be his attempt to take control of the Penguins' lair for revenge. With Skipper having infiltrated the lair and defeated Hans, he was then shipped to the Hoboken Zoo.

He appeared in "The Hoboken Surprise" and helped the penguins defeat the animal androids who had taken over the Hoboken Zoo.

He appeared in "The Return of the Revenge of Doctor Blowhole", working with Doctor Blowhole as part of a plan to take Skipper's memories away.

Dr. Blowhole

Dr. Blowhole (voiced by Neil Patrick Harris) is an evil, mad dolphin scientist and supervillain, who plans to take revenge on the human race: In the past, Blowhole was known as "Flippy", and was forced by humans to perform tricks at Coney Island for their own amusement. Dr. Blowhole is Skipper's arch-nemesis. He only has one (real) eye; his right eye is a robotic one, which hides a scar. He is described by Private and Skipper as being "pure evil, with skin that's surprisingly pleasant to the touch", which Blowhole claims is because he always moisturizes. He appears to love rubbing it in that he has far superior technology than the penguins do, which usually gets on Kowalski's nerves. Since he is a dolphin, he gets around by the use of a Segway-type vehicle while on land, which also projects a glass dome around it and Blowhole to turn into an escape vehicle. He also has a habit of referring to the penguins as "pen-gyoo-wins" (it is implied that he only does this to annoy them). Dr. Blowhole has a habit of having his deep-voiced computer announce the names of his creations in a dramatic voice (provided by Jim Cummings) instead of saying them himself (i.e. Chrome Claw). Unlike most cartoon villains, instead of simply leaving his enemies to die in a deathtrap, he typically stays to oversee their destruction personally, well aware they will likely escape somehow (telling Julien as much when asked), unless he'd be killed as well if he stays (such as locking them in his base as it self-destructs). In the special, he also proved able to hold his own with the combat-trained Skipper in a short physical fight.

Dr. Blowhole is mentioned in the episode "Eclipsed". Blowhole apparently had a plan to extinguish the Sun, as when Private told him that the sun went dark, Skipper believed that Dr. Blowhole had succeeded in his plan, but actually the sun went dark because of a solar eclipse.

In "Roomies", Blowhole sends a walrus spy, named Rhonda a.k.a. Agent 12, to steal Kowalski's "plasma cutter" invention, which acts as a chain saw. The villainous dolphin himself is never seen the episode.

Doctor Blowhole is the main antagonist of The Penguins of Madagascar special Doctor Blowhole's Revenge which also acts as his debut. Dr. Blowhole kidnaps King Julien, believing him to be Skipper's best friend, in an effort to lure the flightless force into a trap so that he can annihilate them once and for all. Julien and Blowhole join forces and captures the "pen-gu-wins". After Blowhole reveals his plan to activate a "Ring of Fire" that he built in the North Pole to flood the world (saying it was inspired by a ring of fire he was forced to jump through when he was in captivity), Mort appears from the lair's skylight and frees the penguins after crashing into a control panel. Blowhole then activates the Ring of Fire after throwing Mort from the control panel. He then sends a giant mutated lobster he devised earlier in the episode called Chrome Claw after Skipper and the penguins (which he'd mentioned he was planning to create during their previous encounter) but they manage to defeat it. After a short battle with Skipper, Blowhole is betrayed by Julien, who claims to have been a double agent the entire time, and the "Ring of Fire" (which the media has dubbed the "Hoop of Heat", much to Blowhole's anger) is deactivated. Blowhole escapes, promising revenge.

Doctor Blowhole was referenced in "Invention Intervention", where Skipper assumed that Kowalski's out-of-control invisible invisibility ray was actually created by Blowhole, commenting that his arch-nemesis had become far more intelligent â€" if harder to follow given the lack of motive behind such an attack â€" since their last confrontation.

In "The Return of the Revenge of Dr. Blowhole", Dr. Blowhole helps Skipper's rival, Hans the Puffin, escape the Hoboken Zoo so he can lure Skipper into a trap. Once he ambushed him, he uses his clown-shaped "Mind Jacker" to steal Skipper's memories so he can use them to infiltrate the penguins' lair so he can turn them into his monster slaves using his "Diaboligizer". When he is about to blast the penguins with the Diaboligizer, he misfires, causing it to hit King Julien's MP3 player infused with Kowalski's experimental power cell (which Julien foolishly thought was a battery). The mix of the three resulted in a giant MP3 monster, which made everyone near it sing uncontrollably. Blowhole soon decides to use the creature to his advantage by winning it over with his beautiful singing voice. Once he was in control, he started using it to destroy New York City. Fortunately, Skipper returned to New York with the help of a spirit guide in the form of Alex the lion as seen earlier in the movie. Skipper distracted Blowhole while the penguins took out the source of the monster's power. With Blowhole foiled again and the monster back to normal, he tried to escape, but Skipper activated the Mind Jacker on Blowhole, wiping out his memories. He was soon returned to Coney Island under his old name of Flippy the Dolphin, where he was forced to jump through the Ring of Fire once again.

He is also mentioned in "The Big S.T.A.N.K", where he was the reason that S.T.A.N.K was created, to lure him to the toilet-shaped stink bomb, but the project was abandoned with the realization that dolphins don't use toilets. He was mentioned again in "Operation: Big Blue Marble" when Skipper suggested Blowhole as a suspect for the recent environmental chaos and random weather conditions that they were experiencing, but Kowalski confirmed that Blowhole was still at Coney Island as Flippy.

In "The Penguin Who Loved Me", it is revealed that Doris the dolphin (Kowalski's love interest) is his sister and his real name is Francis (which Red One, one of his Lobster minions, laughed off). He only created the name Dr. Blowhole since it earned him more respect.

Rhonda

Rhonda (voiced by Kathy Kinney) is a very rude and obnoxious walrus (though she is depicted with buck teeth instead of tusks). She appears in the episode "Roomies" where she is transferred into Marlene's habitat. At first Marlene is excited to have another girl to hang out with, but gets annoyed by Rhonda very quickly due to her disgusting and obnoxious behavior. She asks the penguins to ship her out overnight, but stops the shipment after hearing she is getting taken to a polar bear reserve. Instead they send her to the Hoboken Zoo. At the end, we see that she is one of Dr. Blowhole's agents and stole Kowalski's plasma cutter.

She appears again, as an ally this time, in "The Hoboken Surprise" where she Hans, Clemson, Lulu and Savio help the penguins fight off animal androids in the Hoboken Zoo.

Bada and Bing

Bada and Bing (voiced by John DiMaggio and Kevin Michael Richardson in Brooklyn accents) are giant eastern lowland gorillas. Bada is a gray gorilla and Bing is a brown gorilla. First shown in the "Kingdom Come" episode when they served as thuggish bodyguards for Maurice, they have been seen displaying softer sides upon other occasions, such as when they looked after Mort in "The Penguin Stays in the Picture" while a photographer was there as an unrequested favor to Private, who wanted to be on the cover of the zoo brochure.

The two are attacked by a super-enlarged Mort in the episode "Mort Unbound" in order for him to get King Julien's stolen mango back.

In "Truth Ache," it is revealed that Bing sleeps with a stuffed rabbit named Mr. Bunny.

Burt

Burt (voiced by John DiMaggio in the TV series, Fred Tatasciore in the video games) is an Indian elephant who was just an extra zoo animal during the beginning of the series, but then evolved into a character of greater focus as the series evolved.

He was central to the episode "An Elephant Never Forgets", in which he leaves the zoo to locate a man who had irritated him as a boy by playing his kazoo at the zoo.

In the 2005 videogame version of Madagascar, Burt was called Darnell.

Burt is shown to be obsessed with peanuts, and when he does not eat them, he goes insane as shown in "Jungle Law". He nearly eats Julien when he mistakes him for a peanut.

He played a vital role in the penguins' defeat of the ruthless boa constrictor Savio due to his sheer size preventing Savio from attacking him, squeezing Savio so that the snake's previously-swallowed victims were released.

Roger

Roger (voiced by Richard Kind) is the penguins' American alligator friend who lives in the sewer. They meet him in the episode "Haunted Habitat" when Skipper and Marlene go to investigate in the sewer under Marlene's habitat because of a strange sound. Roger tells the penguins that he is from Florida and that he was the pet of a young boy until he was flushed into the sewers by the boy's parents.

He appears again in "Roger Dodger" when the Sewer Rats terrorize him. Kowalski swapped his mind with Rico's to drive away the Sewer Rats and then had to work to get their minds back into their respectful bodies.

In "Gator Watch," the overload of sewage causes problems for Roger at the time when the penguins were visiting him. The penguins had to work to find the right home for him since the "zoo overlords" would be surprised at the sudden appearance of Roger. First they tried a park with Roger disguised as a log. A woman recognized him. Then the penguins try to disguise Roger as an exhibit at the museum. Two boys ended up giving Roger away. Another idea was to have Roger pose as a gargoyle at the top of a building. Roger fell off the building and into a populated swimming pool. This caused the entire city to end up in a Gator Watch which ended with Roger getting captured by animal control at a theater. When the penguins worked to free Roger, the location Roger was taken to was the Central Park Zoo where an alligator habitat has been set up by the zoo staff.

Since then, he has made background appearances.

Max

Max (voiced by Wayne Knight) is the penguins' stray tabby cat friend who lives in an alley near Central Park. He first meets the four penguins in the episode "Launchtime" when the penguins end up on a rooftop across the street from the zoo instead of on the moon. At first, the penguins thought he was a "mooncat", but at the end they figure out he was just a stray cat. Max is skinny and hopes to catch a bird in his life. He at first wanted to eat the penguins, but was so touched when he was given a can of fish by them that he became their friend instead. He has a very small cameo in "The Helmet".

In "Cat's Cradle", Max tried to escape Officer X. With help from the Penguins, Max and penguins managed to trick Officer X into getting attacked by a wildcat called the Red Rhodesian Slasher.

He appears once more in "Street Smarts" where Max helps the Penguins retrieve Mort from Elmer by teaching them how to be a dog. In the end, Max and Elmer become close friends, but only because Elmer thinks that Max is a lemur.

Duck Family

The Duck Family are a family of ducks that live in a pond in Central Park.

Mother Duck

Mother Duck (voiced by Tara Strong) is a white duck who is the mother of Bradley, Samuel, Ramona, and Eggy.

Eggy

Eggy (voiced by Tara Strong) is a duckling who is the son of Mother Duck and the brother of Ramona, Samuel and Bradley. In "Paternal Egg-Stinct," Marlene found an abandoned egg in her habitat. She gave it to the penguins to look after. Julien wanted the egg for himself. He called it Julien Junior, or JJ for short. Private called the Pony Twilight Sparkle. At the end, Marlene brings Mother Duck to the penguins to get her duckling back.

Unfortunately, the effects of the Penguins' training goes to a point where Mother Duck asks the Penguins to do something to stop him as seen in "Hard Boiled Eggy." Their attempts to show Eggy the dangerous portion of their missions fails, as the duckling goes on a rampage taking down all the animals in the zoo where Eggy inherits 5 traits.

Ramona, Bradley and Samuel

Ramona, Bradley and Samuel are ducklings who are Eggy's siblings and Mother Duck's children. Like Eggy's previous personas before becoming more like Julien in Hard Boiled Eggy, they crave action and mayhem.

Fred

Fred (voiced by Fred Stoller) is a dimwitted squirrel that lives in Central Park and takes everything said literally and has a slow, unemphatic speech pattern. Kowalski met Fred briefly in the "Otter Gone Wild" episode, while the full penguin group met him in "Mask of the Raccoon" where he is friends with Archie despite the fact that Archie is no good.

He dated Marlene in "Otter Things Have Happened", but she broke up with him because what she had thought was him being funny turned out to be just Fred’s ignorance.

In "Field Tripped," Julien recruited him to be the 'smart one' of Julien's all-mammal penguin-team-equivalent.

Fred was part of a treasure hunt in the episode "The Lost Treasure of the Golden Squirrel".

Roy

Roy (voiced by Danny Jacobs) is a short-tempered white rhinoceros, appearing predominantly as an extra zoo animal for the majority of the series. When given more of a role, he often appears to have little-to-no patience for any animal that provokes him, usually responding with threats, such as when a truth-telling Private nearly revealed a secret about him in the "Truth Ache" episode involving a "vacation" in Tijuana.

He plays a pivotal role in the "April Fools" episode when he threatens Julien with retaliation for drawing on his backside with a permanent marker. It is revealed that he was working with the penguins to teach Julien a lesson for his constant April Fools' Day pranks on everyone in August.

In "Antics on Ice", it was revealed that he is a fan of the Lunacorns series due to their positive focus on a horned animal.

Twice in the episode "Action Reaction," he sat on Julien.

Pinky

Pinky (voiced by Kevin Michael Richardson) is a sassy American flamingo, who usually only appears as an extra zoo animal in the series. Her only main role, so far, has been in her debut episode "Go Fish." In this episode she was used for aerial support when the penguins wanted to take fish from a fish delivery truck. Somehow she wound up getting all the fish for herself by the end of the episode.

In the episode "Operation: Good Deed" the penguins need one of Pinky's feathers so that Julien would let Maurice massage Mason. She agrees to give them one after they got her peanuts through a series of additional tasks.

Darla

Darla (voiced by Grey DeLisle in a Western accent) is a very tough Guinea baboon. She appears mainly as a background zoo animal. She was first mentioned in "Assault and Batteries", but made her first appearance in "Out of the Groove". In this episode, she stole Julien's groove after he skunk-bombed her. After several attempts by the Penguins to get the groove back, she finally gives it back when Skipper manages to make Julien apologize.

She had her second major role in "Hair Apparent". In this episode, she began pampering Maurice (thinking he was a male baboon after he got most of his tail fur ripped off, got red paint on his bottom, and got Commissioner McSlade's toupee stuck on his head making Julien jealous. The penguins later take the wig from Maurice to return it, making Darla angry to find out Maurice is a lemur. By the end of the episode, she decides to pamper Mort instead.

Carol and Jillian

Carol and Jillian (voiced by Grey DeLisle) are Darla's baboon friends who are often seen with her.

Shelly

Shelly (voiced by Melissa McCarthy) is an ostrich who lives in the Central Park Zoo. For most of the series, she only appears as a background zoo animal.

Her first major role and her first speaking role so far is in the episode "Love Takes Flightless". In this episode she falls for Rico after he gets her out of cement. She tries to get Rico's attention by hiding his doll in the dumpster. After seeing how upset Rico was about it, she helps the penguins get it back from the garbage truck. At the end, Shelly gets herself a muscular male doll and becomes happy with it.

Becky and Stacy

Becky and Stacy (voiced by Jennette McCurdy and Victoria Justice) are two American badgers that are residents of the zoo. It hasn't been revealed if they are either friends or sisters. They are prone to having to put up with badger stereotypes.

Private was initially fearful of badgers, but Becky and Stacy won his heart by giving him Peanut Butter Winkies. After Private left, Becky and Stacy started badgering Marlene into doing wild and crazy things with them. Marlene, annoyed, tells them to stop badgering her. They ended up attacking Marlene due to the use of badger stereotypes. They were eventually defeated when Marlene fell outside the park and went into her wild state. After that incident, Becky and Stacy appear to respect and fear Marlene since they all have a picnic with the penguins in the park with the two of them being held in between wild Marlene's arms.

Becky and Stacy both make a reappearance in "Tunnel of Love," where they fall in love with the beavers and romance interferes with the beavers' work.

Randy

Randy (voiced by Will Friedle) is a sheep who lives at the Children's Zoo at the Central Park Zoo. Children are constantly coming to pet him. Many of them have sticky hands due to eating too much candy.

In his debut episode "Can't Touch This," one kid in particular hits him and rips his fur until finally Randy cannot stand the torture any longer resulting in Randy biting the kid. The kid screams, causing Alice to come and see what the problem is. The penguins hear the boy's cry as well. When they get there and see the child lie to Alice. Alice apologizes, and puts Randy in a cage. Skipper then says they're going to teach the sheep a lesson for biting a child. Once Randy tells them their story, the Penguins decide to help Randy with his predicament by using various defense methods including: "Repelling" Polymer, Static Electricity, and Hypnosis, all of which backfire. Eventually, Randy ends up back in the Petting Zoo, but as a consultation, Skipper and Rico teach him some self-defense combat techniques to prevent anymore accidents.

Randy appears again in "Operation: Neighbor Swap" where he welcomes the Lemurs into the petting zoo but once Julien gets comfortable, he too grows tired and annoyed of his constant bossing. He helps the penguins formulate a plan to get Julien back to his old habitat.

Muffy, Buffy, and Fluffy

Muffy, Buffy and Fluffy (all three voiced by Kristen Schaal) are three adorable, cute, and cuddly rabbits who reside in the Central Park Children's Zoo. As shown in the show they appear to be cute and innocent but they happen to be very good fighters (evident by how they managed to beat up the penguins with ease).

They made their first appearance in "Operation: Neighbor Swap," when the Lemurs were transferred to the Petting Zoo. When they met the lemurs, they became loyal to King Julien because they heard he was a king (and because of his "magnificent" tail).

Clemson

Clemson (voiced by Larry Miller) is a very descriptive and exuberant bamboo lemur who made his first appearance in the episode "Right Hand Man" as the main antagonist. After being transferred to the Central Park Zoo, Clemson tried to replace Maurice as Julien's right hand man by constantly pampering him. Mort (who had suspected Clemson was evil since he arrived) found out that Clemson was planning to ship Julien to the Hoboken Zoo so that he could take over as king. The Penguins later turn the tables on him by pushing Clemson into the crate to be sent off.

Clemson appeared again in "The Hoboken Surprise" where he helped the Penguins fight off the android animals of the Hoboken Zoo.

Clemson's returns in "King Me" where he informs the lemurs that Julien can't inherite the crown and nearly wins a contest to see who would be king. He was thwarted when the Penguins switched Clemson's brain with Julien's. Julien in Clemson's body then decrees "Julien" king right before they are switched back. Clemson is then chased back to Hoboken by Bada and Bing after Julien insulted them while still in Clemson's body.

Lulu

Lulu (voiced by Jane Leeves) is a female chimpanzee. She first appears in "Monkey Love", in which she was staying at the Central Park Zoo while her habitat in the Hoboken Zoo was being repaired. Once she showed up, Phil fell in love with her. When Mason tries to set the two up, she thinks Mason is asking her out. After several failed attempts by the penguins and King Julien to show off Phil's skills, Phil explains the whole situation to her in Sign Language and she agrees to go out with him before she has to leave.

She appears again in "The Hoboken Surprise", along with Hans, Savio, Clemson and Rhonda, where she helps the penguins fight off the android animals of the Hoboken Zoo.

Doris

Doris (voiced by Calista Flockhart) is a dolphin who Kowalski has a crush on her who is spoken of in several episodes, but as yet unseen. As shown in "Hot Ice," Kowalski sees her as a love interest, though it appears to be unrequited (since Doris wanted someone "hipper").

Doris makes her first appearance in "The Penguin who Loved Me" and is revealed to be the sister of Dr. Blowhole. She wishes for Kowalski to help her free her brother Francis from captivity along with a platypus named Parker. After the penguins and Parker follow Doris to Seaville, they get to Francis' exhibit where it is revealed he is none other than Dr. Blowhole, who has forgotten who he really is and thinks he's Flippy because of his memories being wiped out and is now happily performing tricks for people. Parker than betrays the Penguins and Doris when he reveals he works for Dr. Blowhole and takes him back to the base where all the Lobsters are. As they try to restore his memories, Doris and The Penguins storm in and the aforementioned characters get in a fight. Soon enough Doris accidentally finds Blowhole's Mind Jacker and Parker uses it to restore his memories. Blowhole than captures the Penguins and Doris, and reveals his evil plan is to fire his new weapon "Project Bad Tidings" at the moon, flood the entire world and rule it. But after Kowalski tricks him and they foil his plan and manage to escape before the base explodes. As soon as they get back home, Doris tells Kolwaski that she likes him now and they kiss for the first time.

Savio

Savio (voiced by Nestor Carbonell) is an oversized boa constrictor that has attacked the animals on three occasions. When he was initially transferred to the zoo in "The Big Squeeze", the penguins suspected him of being the cause of the disappearance of the lemurs and Marlene, but his cage appeared inescapable; the only means of departure was a vent that Savio could have used when unfed but would have been unable to access when returning. Skipper realises that he was instead using the vent to access the keypad that controlled his cage locks with his tail, allowing him to open his cage. He was defeated on this occasion when the penguins lured him into Burt's habitat, with Burt throwing Savio around and squeezing him through the bars to expel his previously-eaten victims. Alice later had Savio transferred to the Hoboken Zoo

Savio returned as an enemy in "All Tied Up With a Boa", intending to eat the penguins, the lemurs, and Marlene in revenge for their previous defeat- subsequently intending to regurgitate them and eat them in reverse order to his previous consumption- after immobilising Burt by sending a mouse to the zoo to trick him into running into a wall in panic, but Julien managed to escape by hiding in the popcorn machine and becoming covered in butter, his slippery form resulting in Savio becoming tied up in knots, covered in butter, and sent back to Hoboken.

Savio appeared as an ally in "The Hoboken Surprise" where he assisted the penguins in defeating the android animals of the Hoboken Zoo.

Savio returns again in "The Terror of Madagascar" when Julien has him transferred back to the zoo as he had hired him to eat a baby fossa that he believes is trying to eat him, but the fossa is revealed to see Julien as a father figure and defeats Savio by biting him and chases him back to Hoboken (While annoying Skipper at the idea that the penguins have now not defeated Savio three times & yells in frustration seeing Savio defeated by the fossa instead of them).

The Vesuvius Twins

The Vesuvius Twins (voiced by Atticus Shaffer) are two schoolboys from a very wealthy family. They are disliked by all of the animals of the zoo because of how nasty they are. Even though they are repeatedly banned from going near the animals, they usually get around the ban because of their parent's wealth, e.g. purchasing all of the tickets for an event at the zoo. Private often encourages others to show kindness to the boys, choosing to believe that they are secretly good.

Cecil and Brick

Cecil and Brick (voiced by French Stewart and John DiMaggio) are two criminals that have occasionally run afoul of the Penguins.

Archie

Archie (voiced by Rob Paulsen) is a raccoon with a Robin Hood-like motive and a friend of Fred. He first appears in "Mask of the Raccoon" where he has been going around stealing from the zoo and supposedly giving to the poor. His plot was exposed by the Penguins upon learning about him from Fred and ends up having to give to the poor.

In "The Big Move," Archie appears as a property agent where he tries to sell Rico a house. In this appearance, he is not in his Robin Hood costume and not using a French accent.

Amarillo Kid

The Amarillo Kid (voiced by Jeff Bennett) is a nine-banded armadillo.

A long time ago, the Amarillo Kid challenged Private to a game of mini golf, back when Private was one of the best mini golf players and was known as Mr. Tux. Private quit after the game, which ended in a tie, because he had to hit a little possum girl's ice cream cone (it was a double scoop) to tie it up. Though Private admitted that the pressure made him quit after all. Several years later in "Mr. Tux," he arrived at the Central Park Zoo to challenge Private to a game to prove he's the best. At first, Private refuses to fight him. Enraged by this, the Kid locked down the entire zoo and threatened to blow it up with a thermonuclear reactor. Private battles him, and though the game is close, Private wins with the help of a cricket he had saved early. Ashamed, the Amarillo Kid leaves.

The Amarillo Kid later returns in "Showdown on Fairway 18," but this time has a changed personality. This time, he comes to the Penguins for help when he has lost a mini-golf match against the Gopher Brothers Bo and Gomer and they want his shell (which the Amarillo Kid used in a wager). The Amarillo Kid tells him of the Westchester Putter from Yabogeyhaha, a golden golf putting club which grants its wielder unbeatable power so that even the worst will become the greatest, and he desires to use it in a rematch against the Gophers to save himself. Private and the Amarillo Kid find the Westchester Putter. Through a series of comical foul-ups, they finally claim the Putter. Private hands the Westchester Putter over to the Amarillo Kid on the agreement that he will return it as soon as he has won his rematch. Touched by Private's kindness, the Amarillo Kid confesses his intentions to steal the Putter for the Gophers in order to save his shell as Private's emotions egged him & made him feel guilt for what he was preparing to do. He decides to turn over a new leaf, and voluntarily puts the Putter back, saying he'd rather go shell-less than proving Skipper & Private right that he hadn't changed. Just then, the other penguins and the Gophers show up. The Gophers trap the penguins under a chandelier, take the Westchester Putter for themselves, and escape the building, proving that the Amarillo Kid had been telling the truth and they had tricked Skipper and his team into helping them as Kowalski recapped the gophers' clever ingenuity of their plan. As they make their getaway, the Amarillo Kid stays to free the penguins. The penguins and the Amarillo Kid pursue the Gophers in a mini golf cart chase. Just as it seems the Gophers are going to give the penguins the slip, the Amarillo Kid uses his golfing skills to sabotage their cart and save the Putter. With the Gophers successfully thwarted, the Amarillo Kid thanks Private for believing in him, and they part on good terms... that is, only after Private reminds the Amarillo Kid remembers he is supposed to give the Westchester Putter back.

Miss Perky

Miss Perky (voiced by Tara Strong) is Rico's doll girlfriend.

Granny Squirrel

Granny Squirrel (voiced by Debbie Reynolds) first she appeared as a granny of Fred in "The Lost Treasure of the Golden Squirrel," but then the gang believed she is a ghost.

Barry

Barry (voiced by Kevin McDonald) is a strawberry poison-dart frog who first appears in the episode "Untouchable" where he is a transfer from the Hoboken Zoo and begins terrorizing the animals by temporarily poisoning Skipper, Kowalski and Rico. Private manages to stop him by putting on a steel suit and hugging him, making Barry realize he only needed a friend.

He appears mainly as a background zoo animal in more recent episodes.

Frankie

Frankie (voiced by Jeff Bennett) is a homing pigeon who first appeared on "Launchtime."

The Red Squirrel

The Red Squirrel (voiced by Jeff Bennett) is an evil red squirrel who appears in the episodes "The Red Squirrel," "A Visit From Uncle Nigel," "Our Man in Grrfurjiclestan," and "Nuts to You."

The Broches

The Broches are the trio of cockroaches who appears in the episode "Stop Bugging Me."

Mike

Mike (voiced by Zand Broumand) is a member of the Broches.

Duane

Duane (voiced by Brian Posehn) is a member of the Broches.

Eddie

Eddie (voiced by Jerry Trainor) is a member of the Broches.

Frances Alberta

Frances Alberta (voiced by Megan Hilty) is the presumptive head zookeeper at the Hoboken Zoo who appeared in the episode "The Hoboken Surprise." Frances was first shown to be a cheerful zookeeper who had worked to clean up the zoo's poor conditions, gaining the respect of resident animals Rhonda, Hans, Clemson, Savio, and Lulu. Frances also afforded the animals under her care many amenities, such as fine cuisine, massage chairs, and a fountain of cheese. It is later revealed that Frances never had the respect of her animals, whom she had simply sealed away under the ground and replaced with android look-alikes, all in an effort to meet her demanding standard for cleanliness and to get the attention of Parks Commissioner Pervis McSlade. At the end of the episode, Commissioner McSlade fires her and the Hoboken Police arrests her for making android look-alike animals and assault of a Bartender.

Kitka

Kitka (voiced by Kari Wahlgren) is a female peregrine falcon whom Skipper falls in love with. She only appears in "The Falcon and the Snow Job".

Nurse Shawna W. Smith

Shawna W. Smith (voiced by Joanna Garcia) is a zoo nurse who only appears in the episode "Love Hurts".

Special Agent Buck Rockgut

Special Agent Buck Rockgut (voiced by Clancy Brown) is a rockhopper penguin who first appears in "The Red Squirrel".

Hunter

Hunter (voiced by Ciara Bravo) is a young female leopard seal who doesn't eat penguins. She only appears in the episode "Operation: Antarctica".

Uncle Nigel

Uncle Nigel (voiced by Peter Capaldi) is Private's uncle, who works as a spy for MI6, and comes to New York to visit his nephew and to find The Red Squirrel. His first, and so far only, appearance is in "A Visit From Uncle Nigel."

When around the penguins (minus Private), Nigel acted like a "posh nancycat". This was all part of his cover story. When captured by The Red Squirrel, Nigel was forced to tell Skipper, Kowalski, and Rico that he actually was a spy.

It is unknown if Nigel really is Private's uncle in blood or not (due to being a different species of penguin). Though it is possible that he is either an honorary uncle or perhaps one of Private's parents was the same species of penguin Nigel is and he is their brother whilst the other parent was the same species as Private.

Manfredi and Johnson

Manfredi and Johnson (voiced by James Patrick Stuart and Danny Jacobs) are penguins who were former members of Skipper's team. They are typically cited for the nature of their demise or a mistake they made as an example of what can happen if a penguin lets their guard down or neglects their training. The exact method of their demise varies in each instance and in some cases are contradictory: killed by an exploding trap, killed by flying piranhas, losing vital internal organs, etc.

They are finally seen in "The Penguin Who Loved Me", in an enclosure in Seaville both with bruises and injuries, but very much alive. However, they have somehow injured their eyes. Manfredi (the larger sized penguin) has lost his left foot now replaced by a stick while Johnson (the Kowalski-sized penguin) broke his right flipper and seems to have a burn on his body. Johnson noticed the penguins and Doris "flying out" of Seaville and banged on the glass panel for help. Unfortunately for them, the penguins never noticed their desperate pleas and swam away saying they were never going to come back ever again.

Strangely they do not appear in Penguins of Madagascar, implying that they may have met the four at a much later time.

Characters in All Hail King Julien

Clover

Clover (voiced by India de Beaufort) is a green-eyed crowned lemur who is King Julien's bodyguard. Her personality is similar to Skipper and Kowalski.

After the events of the first episode, Clover becomes his commander. She is the main protector King Julien (since the other guards ran away) Despite the fact that she thinks the new king isn't very bright she is fanatically loyal to him and is even willing to die for her king. Unlike his uncle, Julien treats her with a lot more respect (for example at the end of episode one he points out that he noticed she was the only one of the guards who didn't run away and because of that his promoting her to commander). Her shocked response to the comment implies that uncle Julien never bothered to even compliment her. A running gag is that she acts like she is talking with someone through a radio earpiece.

When in the second episode, one of the lemur's points out that she doesn't even have an earpiece where she applies very painfull pressure to the lemur's wrist and once again talking through her imaginary earpiece says she's got a troublemaker and that she's bringing him in for interrogation. She is obsessed with order and new things, and she knows at least 45 arts on how to kill a man with just her teeth. Her eyes seem to flame fire sometimes. She calculate math in less than 10 seconds. She is able to take out one fossa but can't take out a whole herd.

She didn't believe in having a fun life until she slid down the water slide going after Mort during Uncle King Julien XII's return. Clover still stuck with King Julien over his uncle.

In "Eat Pray Shove", King Julien tells Clover to take a vacation, with Xixi and Mort. Xixi and Mort seem to enjoy, while Clover doesn't. The two discover a book and it is revealed that Clover writes about a huge guy. She leaves and she meets a muscular and handsome lemur called the Sage Moondancer.

In "He Blinded Me with Science," it is revealed that Clover writes fan-fiction. She also makes sure that Masikura keeps it a secret from the other lemurs or else.

In "Crimson and Clover," it is revealed that Clover has a twin sister named Crimson who is the opposite of her.

Masikura

Masikura (voiced by Debra Wilson) is a psychic chameleon who appears once you call her. She sticks her tongue on your head and analyzes you for predictions or reading thoughts. He told Julien's uncle that the king of the lemurs would be eaten by the Foosa. With that, Uncle King Julien hands over the crown to his nephew and leaves. At the coronation, the Foosa interrupted and take some lemurs with them. Masikura told King Julien, Maurice, and Clover about what happened. King Julien goes to the Foosa territory to save the lemurs, he got bitten on the butt, he shouts and the boulders fall down, scaring the Foosa. Julien was relieved that Masikura was wrong about the Foosa eating him. Masikura appears and say she didn't say the Foosa wouldn't eat all of him completely and points at his butt.

In "Poll Position", King Julien calls Masikura to use her tongue on Xixi's head to find out who's the one hater. Masikura denined, but King Julien grabbed her and starts shooting her tongue at Xixi. Her tongue hits Maurice's head and predicted grape jelly, a funnel, and a pair of woman shoes. Maurice said it was spring break.

In "He Blinded Me with Science," Masikura played a big role, where she helps King Julien dance for the sky god Kevin to make it rain. When King Julien meets Timo, a tenrec scientist, he automates everything in the kingdom and they got hooked up on science. Later, King Julien let Masikura go, much to her dismay. She meets Mort in his tree stump and insisted to stay with him. During her stay, Masikura gets creeped out with Mort, saying he "sleep hugs" and says that he is a girl. Then, a fire breaks out from the generator, King Julien apologizes to Masikura and he, Maurice, and Clover ask for her help. Masikura believes that if they combined science and the rain that would put out the fire and save the kingdom. Timo makes a makeshift hose and King Julien danced on it to get the water out, thus putting the fire out.

Prince Barty

Prince Barty (voiced by John Michael Higgins) is a ring-tailed lemur who is the father of King Julien XIII and the husband of Princess Julienne.

Princess Julienne

Princess Julienne (voiced by Anjelica Huston) is a ring-tailed lemur who is the mother of King Julien XIII and the wife of Prince Barty.

Uncle King Julien XII

Uncle King Julien XII (voiced by Henry Winkler) is a ring-tailed lemur who is the uncle of King Julien XIII and is one of the antagonists of the series. As King Julien XIII's predecessor, King Julien XII is a lazy, strict, paunchy, cowardly, and older lemur oozing smarminess where he had strict rules that kept the lemur kingdom quiet in order to keep the Fossa away. In the first episode, he gives the crown to his nephew, Julien and leaves when he learns from Masikura that the king of the lemurs will be eaten by the Foosa. When King Julien saves the captured lemurs, he got bitten on the butt and is still alive. When King Julien XIII thrives as king, his devious uncle will do any conniving thing to regain the throne.

Uncle King Julien returned in the fifth episode, not happy that his nephew is still alive and set to reclaim the throne from him. When King Julien was sliding down the water slide, his uncle turns off the water, leaving his rear end on fire. King Julien then sees his uncle and has a hard time remembering him, much to his annoyance. Maurice helped Julien remember by placing his crown on his uncle's head. He made up a story and tricked his nephew that he heard the Fossa were planning an attack and Julien volunteers to go to their territory and find out with Maurice in a Fossa disguise. During his absence, Uncle King Julien made changes to his throne room. Mort goes up to the throne room and sees Julien's uncle. Uncle Julien takes a seriously hatred of Mort. Mort says that Julien, (the other Julien), is the one and only king and is "a billion times better than him", which makes him mad. Uncle Julien orders Clover to destroy Mort, but Mort says he'd rather be destroyed by "his King Julien" and runs off crying. After seeing the changes his nephew made, he orders Clover to close down everything Julien put up. When Clover closed everything down, she chased Mort down the water slide when he was violating the rule against it and she actually enjoyed it. Clover tells Julien's uncle what happened and says the kingdom is happy for the first time in a long time and vows to help improve when Julien comes back. When Julien's uncle says that Julien is not coming back, Clover, discovering Julien's uncle's plot, angrily forces Uncle Julien to say it again and drags him to the Foosa territory. They see Julien and Maurice in the Fossa suit surrounded by the Fossa. Uncle Julien is unhappy that Clover wants his nephew as king. Before Uncle Julien can hit Clover, she hits Uncle Julien and uses him as bait to attract the Foosa and saves Julien and Maurice, while fighting the Fossa via suit. Julien knows his uncle's plot to reclaim the throne and is glad that Clover chose him, even though she didn't think the kingdom should be fun 24/7 but knows living in fear is worst. While taunting the Fossa with Uncle King Julien still as bait, Maurice wondered about what will they do with him, as he is not to be trusted. Julien says that his uncle is still family and "family always deserves a second chance." He suggests that they should give his uncle "a chance at a new life." The episode ends with Uncle Julien in the Foosa suit that is being cuddled by a Fossa girl much to his dismay. In a later episode he creates his own kingdom, Fearland, tricks his nephew into giving him all his subjects, and imprisons them there as "citizens"; he is later imprisoned alone inside his own little dystopia.

He shows no love or care to his nephew when he tricks him into being king and going into the Fossa territory, leaving him to be eaten.

According to Masikura in "Viva Mort," Julien's uncle is the worst king because he did some things to other kingdoms and never apologize. Masikura told King Julien he doesn't have to be his uncle, when she is trying to help him apologize to Maurice.

Ted

Ted (voiced by Andy Richter) is a Golden bamboo lemur who makes appearances in the series. He makes appearances in every episode, and played big roles in "Eat Prey Shove," and, "Diapers are the New Black." He is very gentle, nice, and speaks with a valley girl accent. He was/is Clover's assistant captain of the Ringtail guard.

In "Poll Position" when people disliked King Julien, Ted would eat the kumquats off the pole, decreasing the popularity of people liking King Julien. He states it was his dream job and it has become a nightmare, when he ate too many kumquats because more people no longer like him. Maurice ask Julien why does he have to eat the kumquats and Julien answered it's the "only thing that brings him joy."

In "Eat Prey Shove," Ted once covered for Clover when she was sent on a vacation with Mort and Xixi. Ted unknowingly caused the Fossa to take advantage of Clover's vacation to attack King Julien's kingdom.

Willy

Willy (voiced by Cody Cameron in Madagascar, Jeff Bennett in All Hail King Julien) is a lemur who first appears in the first episode as one of the captured lemurs. King Julien saves them from the Foosa. He appears in the second episode in which he is told that he is to become the captain of King Julien's first navy ship named Old Porceleinesides. He appears to be slightly smarter than the other lemurs as he was immediately distraught by the fact that the ship he was to captain didn't look very seaworthy. Still with some motivation from Maurice (who poked him with a stick to get him to move towards the boat) he was smart enough to accept the honor. His worries about the boat being unseaworthy are confirmed later in the episode when the boat starts sinking, a few minutes later after King Julien flipped the switch to release the boat into the water. He is later seen in the episode being attacked by very aggressive dolphins while the boat is sinking. When he is angry with King Julien for putting him in this predicament (the sinking ship) he lets himself be bribed with the mango's King Julien was sharing to increase his popularity. After catching the mango he screams, "You're the best, King J!" He is either very lucky, able to swim or rescued from the water by Clover, as he appears in the next episode in which he celebrates Franksgiving in the honor of the sky god named Frank. He was trying to smuggle in more than one wishing stone in the wishing balloon something which he is called out for by Maurice.

Willy was briefly seen in the first film when he was called out during King Julien's meeting about what to do with Alex, Marty, Melman, and Gloria. He states that he likes them.

Xixi

Xixi (voiced by Betsy Sodaro) is a very trustworthy toucan in news who informs the lemur kingdom on current events. According to King Julien, Xixi also a very trustworthy Toucan in friendship. She covered King Julien's coronation party but fled when the party was raided by the Foosa in the first episode.

She appears again in "Poll Position" in which she reports that 99% of the kingdom thinks the new king (Julien) is doing a good job. This report sets both episode plots in motion namely King Julien attempting to locate and turn the hater while also allowing Clover to launch an investigation to find the hater and make sure he isn't trying to recruit more lemurs to his side to cause an uprising.

She makes another appearance in "Eat Pray Shove", when she happily accepts to go on a vacation with Clover and Mort. Xixi and Mort enjoy relaxing on the beach, while Clover doesn't. Xixi and Mort have a fun time while Clover went off and meet a handsome and muscular lemur.

Hector

Hector (voiced by Jeff Bennett) is an incredibly moody lemur. According to Maurice, Hector hates everybody and everything. He is also a bit odd as for example when Clover asked if he didn't want to see King Julien to help him solve his problems, he replies to her that he likes his problems. According to Xixi, he has a very large family. Despite his bitterness, he does admit that King Julien is doing a better job than the previous kings. Hector also adds that he thinks that it is not a great accomplishment as he thinks the previous kings were much more incompetent than Julien was.

Horst

Horst (voiced by Jeff Bennett) is a blue-eyed black lemur. As said by Clover in the episode My Fair Foosa, not much is known about him. Kind of a loner, fears change, often seen holding a beverage. He is also known as Dj Glitter-Bunz. Which is Revealed in All Hail King Julien Episode: King Julien Superstar.

Dorothy

Dorothy (voiced by Sarah Thyre) is a mongoose lemur who is the wife of Ted.

Pancho

Pancho (voiced by Danny Jacobs) is a crowned lemur. He is mentioned in "Poll Position" when Xixi mentions him in her report stating that he is looking cozy for he's canoodling with a new lemur lady and questions if love is in the air. King Julien agrees that he hopes so and states that "Pancho needs it."

Sage Moondancer

Sage Moondancer (voiced by Jeff Bennett) is a buff and spiritualistic lemur.

Tammy

Tammy (voiced by Debra Wilson) is a common brown lemur who is the mother of Todd. While she loves her son, Tammy tends to pressure him.

Todd

Todd (voiced by Kevin Michael Richardson) is a common brown lemur who is the son of Tammy. he is a child prodigy.

Timo

Timo (voiced by David Krumholtz) is a tenrec scientist who is an expert on technology.

King Julien first encountered Timo in "He Blinded Me with Science" where he replaced the batteries in King Julien's boom box after it ran out of power during King Julien's dance to get the Sky God Kevin to bring rain to his kingdom. Upon introducing Timo to the lemurs, King Julien had them receive the technology that was salvaged from Timo's area. The lemurs get hooked on the technology until a power surge happened that destroyed Timo's generator and started a fire. With help from Masikura, King Julien, Maurice, Mort, Clover, and Timo were able to find ground water to help stop the fire.

In "Viva Mort," Timo invented a device that would help King Julien to make him embarrassed as a way to make it up to Maurice. Due to the device not being plugged in, Maurice was put through the device and embarrassed instead.

In "The Really Really Big Lie," King Julien asks Timo to build a giant mechanical gecko to make it believable to cover his lie. The robot gecko went on a rampage, leaving the kingdom in a panic. Timo says that the off switch is on the bunny ears because Todd loves bunnies. King Julien, Maurice, and Clover managed to shut down the mechanical gecko and escaped from a fall.

In "One More Cup," King Julien, Maurice, and Mort find a bag of coffee beans and take it to Timo, who made a coffee machine. King Julien drinks the coffee and feels the energy. King Julien and Timo sells the coffee to the lemurs that Julien calls "Brown Julien".

In King Julien Superstar, Timo helped King Julien and the gang become famous superstars by "tuning" their voices to sound good.

King Joey

King Joey (voiced by Bill Fagerbakke) is a jumping rat who is the ruler of Madagascar's jumping rats. Like his kind, King Joey is not very bright.

Becca

Becca (voiced by Sarah Thyre in a Southern accent) is a red/orange-eyed black lemur who is Abner's wife. Becca and Abner encountered King Julien in his alias of "Banana Guy Mike" where he had helped them fix their watering hole. She and her husband vowed to get King Julien off his throne to avenge the "death" of Banana Guy Mike.

In "Viva Mort," Becca and Abner have started the Lemur Alliance Liberation Army (LALA) to get revenge on King Julien. They duped Mort into helping them until their movement was discovered by Clover. Though Becca and Abner are thwarted, Clover had to leave to keep King Julien from getting stung by a scorpion that was placed in a package that they had Mort give to him.

Abner

Abner (voiced by Diedrich Bader in a Southern accent) is a blue-eyed lemur who is Becca's husband and is not very bright. Abner and Becca encountered King Julien in his alias of "Banana Guy Mike" where he had helped them with their powers. He and his wife vowed to get King Julien off his throne to avenge the "death" of Banana Guy Mike.

In "Viva Mort," Abner and Becca have started the Lemur Alliance Liberation Army (LALA) to get revenge on King Julien. They duped Mort into helping them until their movement was discovered by Clover. Though Abner and Becca are thwarted, Clover had to leave to keep King Julien from getting stung by a scorpion that was placed in a package that they had Mort give to him.

Karl

Karl (voiced by Dwight Schultz) is a devious fanaloka and is one of the antagonists of the series. Part of his history mentioned that he has a brother named Bruce who was better at things than him, especially making coffee and Karl makes "poo coffee". His catchphrase is "not yet at least".

After he did some attempts on King Julien's life where Mort took a blow to every attempt, Karl makes himself known to King Julien and invites him to his lair with the directs on the brochure. After Clover and Maurice are captured, King Julien makes his way to Karl's lair at a crashed zeppelin where Karl mentions his plans to raise the zeppelin and take over the world with King Julien as his partner. While Karl was ranting about his plans, King Julien manages to free Clover and Maurice. When King Julien, Clover, and Maurice exit the zeppelin, they are confronted by Karl who prepares to fire a laser at him until the balloon carrying the rocks with every wishes that the lemurs wrote on them for the Sky God member Frank falls from the sky and lands on Karl. Seeing his "Surprise Me" wish roll against him, King Julien took it as a sign that Frank had helped him defeat Karl.

In "One More Cup," it is revealed that Karl survived and returned where he hooked the whole kingdom on caffeine which was part of his plan. He states he watches King Julien fooling around in the Cove of Wonders instead of following his kingly duties. Karl threw a bag of coffee beans in the ocean and knew that Julien would find it because it has the word "king" on it. The hyper kingdom threw out King Julien, Maurice, and Clover. With the help of Mort (who became intelligent from the coffee), they plan to destroy Karl's coffee supply. They approach to Karl's zeppelin while Smart Mort walks through the fields where he meets jumping rats working there. Karl returns, finding them, and tries to destroy them when the jumping rats break through the door demanding a raise from their boss, even the ones cornering Julien, Maurice, and Clover. Later, Karl surrounds the group and prepares to fire the laser, from earlier at Julien until Mort bites him on the foot. King Julien has the weapon and sees that Karl has Mort in his clutches and says that he has one shot, him or Mort. Julien, having no choice, aims the laser at the fields destroying the crops leaving Karl devastated worrying what will Bruce will say when he returns from vacation. King Julien now aims the laser at Karl who states that it charges. Karl escapes in a hot balloon with Chauncey driving it. Karl vows that he will return and will never stop getting rid of King Julien.

In "Are You There Frank? It's Me, Julien," Karl creates a special armor that allows him to pose as Frank the Sky God in order to get the Lemurs to oppose Frank's brother Larry the Volcano God by sacrificing King Julien. With help from Mort operating the Larry armor that Timo built, King Julien repels Karl. in "revenge of the prom" Karl reveals that he was in all of Julien's old high school classes, but julien never noticed him.

Chauncey

Chauncey (vocal effects provided by Kevin Michael Richardson) is a Madagascar hissing cockroach who is Karl's pet and henchman.

Crimson

Crimson (voiced by India de Beaufort) a red-furred crowned lemur who is the twin sister of Clover and is the opposite of her.

Rob McTodd

Rob McTodd (voiced by David Koechner) is a lemur who was King Julien's old friend.

Vigman Wildebeests

Vigman Wildebeests(voiced by Jeff Bennett) is a cocky blue wildebeest, he was a rival of Clover's grandmother. In the episode 'Eye of Clover", he enters the tournament and defeats every players. However he is defeated by Clover and later has a rematch against Clover's Grandmother.

Dr. S

Dr. S (voiced by Jeff Bennett) is a cobra who works as an self-taught underworld doctor. Though it takes work to get a doctor's license, Dr. S's claim is that he can't get busted if he practices medicine in his own cave.

Fred the Giant Scorpion

Fred the Giant Scorpion (voiced by H. Jon Benjamin) is a giant scorpion that lives in the desert parts of Madagascar.

Captain Ethan

Captain Ethan (voiced by Rob Paulsen) is a rat who is the leader of the pirates that operate in the waters of Madagascar.

Merry Madagascar characters

Donner

Donner (voiced by Jim Cummings) is the leader of Santa Claus' reindeer. His name is revealed in "Santa Claus is Coming To Madagascar".

Cupid

Cupid (voiced by Nina Dobrev) is one of Santa Claus' reindeer who falls in love with Private.

Reception

Sean Axmaker of Seattle Post-Intelligencer praised the characters, noting the "marvelous character animation" and "the palpable camaraderie between animal buddies" kept Madagascar: Escape 2 Africa "rolling". Axmaker also commended the voice of Zuba, played by Bernie Mac, saying "his vocal makeover is so complete that you may not recognize his voice, but you will appreciate the warmth and vivid personality of his creation."

References